Highlights from 2001
Updated: 4 October 2006

Smithsonian Institution
National Museum of Natural History

Research Training Program

Main Page
2001


26 May 2001 - 4 August 2001

A total of 18 students were selected to participate in the 2001 session of the Research Training Program including 3 international students representing Brazil, Canada, and Yugoslavia.

Schedule of Events  |  Poster  |  Program Summary
Student Abstracts
  |  Photo Gallery
The Ethics Discussion

Applicant Pool

7,500 recruitment posters were distributed to worldwide locations.

Over 5,000 information requests were recorded.

219 applications were received.

18 students were selected to participate.

Semi-finalist list

Finalist list

Participants

Schedule Summary

Apartment Assignments

Recruitment Poster


Research Training Program
Project Summaries

(Click on the name below to go to the individual project summary page.)


Meet the 2001 participants
and read about their research projects
2001 RTP Group Photo


Back row (left to right)
Katarina Topalov, Anastasia "Stacie" Poulos, Matt Friedman, Michael Holcomb, Trevor Krabbenhoft, Jeff Saarela, Laura Holladay, Yana Reid, Alexandre Ribeiro.

Front row (left to right)
Sharon Wilson, Dawn Stodden, Erin DiMaggio, Ainsley Seago, Marita Davison, Abigail "Abby" Knee, Yolanda Chacon, Courtney Porreca, Shannon Schwaller, and Julianne Rowehl.

In addition to our research reports, be sure to check out our personal summaries about our summer at the Smithsonian.


Yolanda Chacon

  • Gender: Female.
  • Ethnicity/Race: Hispanic.
  • Institution: New Mexico State University.
  • Status: Sophomore.
  • Major: Biology.

Career Goals: After graduate school achieve a career in either Ornithology or Medicine.

Funding: Participation in the 2001 summer session of the Research Training Program was supported by the National Science Foundation Research Experience for Undergraduates program, Award: DBI-9820303.


Update - December 2002: Yolanda is finishing an undergraduate degree at New Mexico State University. She recently presented a poster explaining the results of her RTP research at The American Biomedical Research Conference for Minority Students. She plans to attend graduate school and maybe even study birds - much to the delight of her RTP research advisor Carla Dove!

Yolanda J. Chacon

Project Advisor: Dr. Carla J. Dove, Department of Systematic Biology, Division of Birds.

Project Title: Microscopic feather character analysis of the cranes, rails and allies (Avian Order: Gruiformes).

Research Abstract: Twenty-six feather characters (both micro- and macroscopic) of 12 species of birds within the avian order Gruiformes (cranes, rails and allies) were described, photographed and then investigated using parsimony analysis to determine if feather characters in this group of birds provide phylogenetic signal.

Feather characters were analyzed separately and in combination with mitochondrial 12S rDNA data (Houde, MS in prep) and resulting trees were compared. Strict consensus of two trees obtained from feather characters (TL= 91, CI= 0.68, RI= 0.59, RC= 0.40) shows that these characters have indices comparable to the tree obtained from molecular characters (TL= 2147, CI= 0.53, RI= 0.38, RC= 0.20). However, the resulting feather and DNA trees agree only in some terminal taxa relationships.

Although the rails (Rallus, Laterallus) are sister to each other according to both datasets, the DNA characters place Sungrebe (Heliornis) as the closest relative to rails whereas the feather characters place Mesitornis (roatelo) as the sister to that group. The strict consensus feather trees and the DNA tree are congruent in the relationship of Cariama (seriema) to Choriotis (bustard). Grus (crane) and Aramus (Limpkin) are in the same clade according to both trees, however feather characters place Heliornis within that clade instead of Psophia (trumpeter) as is suggested by DNA characters. The DNA tree does not resolve the status of Mesitornis but rather places this taxon as an outgroup to the main clade. A single tree of all combined characters resulted in a tree with indices of TL= 2086, CI=0.54, RI= 0.42, RC=0.23 and was more similar to the tree obtained from the DNA characters.

Although more research is needed on the feather characters in this group of birds, this study provides evidence that feather characters produce trees that cluster into 'natural groupings' in parsimony analysis and supports the idea that these characters can be useful for feather identification and phylogenetic analysis.

Publication: Chacon, Y.J, and Dove, C.J, In Prep. (To be submitted in summer 2002) Microscopic feather character analysis of the cranes, rails and allies (Order: Gruiformes). Proposed for publication in the Wilson Bulletin.

Project Summary  |  Letter of Gratitude



Erin DiMaggio

  • Gender: Female
  • Ethnicity/Race: White
  • Institution: University of Michigan.
  • Status: Freshman.
  • Major: Geology.

Career Goals: After obtaining a Ph.D. in Geological Sciences, pursue a career as a volcanologist.

Funding: Participation in the 2001 summer session of the Research Training Program was supported by the National Science Foundation Research Experience for Undergraduates program, Award: DBI-9820303.


Update - December 2002: Erin is still an undergraduate student at the University of Michigan. She spent the summer of '02 serving an internship with NASA at the Jet Propulsion Lab in Pasadena, California, specifically focused on the Martian Exploration Program with Arwen Vidal (1999).


Erin N. DiMaggio

Project Advisor: Dr. Sorena S. Sorensen, Department of Mineral Sciences.

Project Title: Evidence for Feldspar Replacement in Altered Metamorphic Tuffs in the Duck Lake Volcanic Suite

Research Abstract: Metavolcanic rocks can be powerful tools for documenting the complex histories of multiple episodes of fluid-rock interaction in long-lived arc crust. The Duck Lake area of the Mammoth Crest, eastern California, is part of a lithotectonic assemblage exposed along the eastern front of the Sierra Nevada. Meta-rhyolite tuffs at Duck Lake show complex K, Na, and Ca systematics and disequilibrium feldspar phase relations that reflect a superposed history of contrasting metasomatic systems.

Using cathodoluminescence and mineral and whole rock compositions, a reconstruction of the metasomatic system's history was prepared. Cathodoluminescence petrography shows igneous phenocryst and ground mass feldspar completely replaced by near-end-member orthoclase (Or>95 after sanidine) and or albite (Ab>95 after plagioclase).

These textures resemble those of nearby meta-tuffs in the Ritter Range Pendent (on strike, 20 km NW) and of unmetamorphosed Tertiary tuffs (Creede, CO, Socorro, NM) that have undergone low temperature K- or K/Na- metasomatism. In those rocks alkali alteration occurred soon after deposition. These early features are overprinted by Ca-metasomatism that is manifested by calcite veins and grossular + epidote veins with calcic plagioclase (An80-100) vein envelopes. The early pseudomorphs of phenocrysts and groundmass feldspar are replaced by calcic plagioclase in this event. Whole-rock geochemistry shows concomitant gains of Ca and Sr and loss of Na. CaO/Na2O for tuff samples ranges to 13, a value >6 times that of unaltered ashflow tuffs. K/Na values for Duck Lake tuffs are similar to those seen in the Ritter Range, Creede, and Socorro, but display distinct slopes on plots of K/Na versus Rb/Sr.

Textures indicate that Ca was sourced by hydrothermal fluid derived from carbonates, probably locally. The Ca-rich silicate veins cut metamorphic fabrics that Tobisch et al. (2000) regionally attributed to batholith emplacement, indicating Ca-metasomatism occurred during retrograde phases of contact metamorphism. Evidently, as seen in many ore deposits, cooling contact metamorphic systems can be accompanied by large amounts of mass transfer.

Project Summary  |  Letter of Gratitude



Matt Friedman
  • Gender: Male.
  • Ethnicity/Race: White.
  • Institution: University of Rochester.
  • Status: Junior.
  • Major: Biology and Geology.

Career Goals: After obtaining a Ph.D. in paleontology, achieve a career as a university professor conducting research on extinct and extant fishes.

Funding: Participation in the 2001 summer session of the Research Training Program was supported by the National Science Foundation Research Experience for Undergraduates program, Award: DBI-9820303.


Matthew S. Friedman

Project Advisors: Dr. James C. Tyler and Dr. G. David Johnson, Department of Systematic Biology, Division of Fishes.

Project Title: A putative lampridiform from the Oligocene of Peru.

Research Abstract: A nearly complete
fish skull and two associated vertebrae (USNM 494403) from the Mancora Formation (Oligocene) of Peru is tentatively identified as a lampridiform and its osteology is described.

Though no synapomorphic characters are observed that definitively place the specimen within the order Lampridiformes, features of the anterior portions of the skull suggest that a lampridiform synapomorphy may have been present in life. This feature, combined with numerous similarities of uncertain polarity found in both the fossil and recognized lampridiform taxa, suggests that cautious placement of the specimen within the order Lampridiformes is warranted.

Other characters in the fossil, notably the position of the foramen magnum relative to the exoccipital condyles, presence of a supraoccipital crest, and a well-developed first vertebra, exclude the 5 taeniosomous families of higher lampridiforms as well as the bathysomous Lamprididae, allowing for placement of the specimen within the Veliferoidei, the most basal suborder.

This specimen is the only fossil lampridiform known from South America. Given the pelagic habit of all extant members of the Lampridiformes, as well as a billfish fossil from the same locality, it seems likely that the Mancora formation was deposited in a deep-water, offshore environment.

Project Summary  |  Letter of Gratitude



Mike Holcomb

  • Gender: Male.
  • Ethnicity/Race: Native American.
  • Institution: University of Idaho.
  • Status: Sophomore.
  • Major: Biology and Chemistry.

Career Goals: After obtaining a Ph.D., achieve a position in an academic institution conducting research and courses on coral reef biology.

Funding: Participation in the 2001 summer session of the Research Training Program was supported by the National Science Foundation Research Experience for Undergraduates program, Award: DBI-9820303.


Michael C. Holcomb

Project Advisors: Dr. John Pandolfi and Dr. Ian
Macintyre, Department of Paleobiology.

Project title: Use of X-radiographs for coral identification.

Research Abstract: Recent work has shown the Montastrea annularis species complex to consist of at least three extant species. These species can be distinguished using several techniques, however, in death assemblages and fossilized material, identification using conventional techniques is problematic. Many of the characteristics commonly used depend upon having whole colonies, or unabraded corallite surfaces.

Characteristics measured using X-radiographs were investigated for their potential to distinguish both modern and fossil members of the M. annularis complex. Characteristics showing significant differences among the species included: maximum corallite growth angle, band intensity, growth rate, fan length, corallite angle along the growth axis, budding angles, and colony curvature. These characteristics are much less prone to loss due to taphonomic processes than are many more conventional ones.

These results suggest that X-radiographs may serve as useful tools for separating members of the M. annularis complex. Considerable geographic and/or temporal variation was noted as well as interspecific variation.

Publication: Holcomb, M., Pandolfi, J.M., Macintyre, I.G., and Budd, A.F. In prep. Potential use of X-radiographs to distinguish members of the Montastraea annularis species complex. Proposed for publication in Coral Reefs.

Project Summary  |  Letter of Gratitude



Laura Holliday

  • Gender: Female.
  • Ethnicity/Race: White
  • Institution: The University of Michigan
  • Status: Recent graduate
  • Major: Geological Sciences

Career Goals: After obtaining a Ph.D., become a professor or researcher at an academic institution.

Funding: This student's participation in the summer 2001 Research Training Program was supported by a grant from the Smithsonian Women's Committee to provide an opportunity for international students and graduating seniors to participate in the program.


Holladay, Laura:

Project Advisor: Dr. Brian Huber, Department of Paleobiology.

Project Title: Ontogenetic morphometric comparison of middle-late Eocene pseudohastigerinid planktonic foraminifera using X-radiograph image analysis

Research Abstract: Middle to late Eocene species of the planispiral planktonic foraminiferal genus Pseudohastigerina include P. micra and P. naguewichensis, which are widely recognized and easily distinguished species. Less readily distinguished are Pseudohastigerina acutimarginata, which resembles P. micra but has a more pinched peripheral margin, and a previously unrecognized morphotype that resembles P. micra in its early ontogeny but becomes more evolute later in its ontogeny.

To test whether or not these latter forms should be synonymized under P. micra, we obtained high resolution X-radiograph images of specimens mounted in axial and edge views to characterize ontogenetic changes in shell and chamber size and shell coiling. Populations of Pseudohastigerina were selected from a geographic array of sites and from several stratigraphic levels within the middle and upper Eocene. The biometric analyses demonstrate significant differences in chamber growth trajectories, chamber coiling, maximum test size, and roundness of the ultimate chamber face between populations of P. naguewichensis and the other three pseudohastigerinid morphotypes. Populations assigned to P. micra and P. acutimarginata are statistically indistinguishable in all coiling and size measurements, and comparison of ultimate chamber roundness shows a continuous range of circular to peripherally elongate shapes. The strongly evolute pseudohastigerinids are similar to P. micra in all measured features except final chamber number and maximum test size.

We conclude that P. micra and P. acutimarginata are synonymous and probably represent ecophenotypes, whereas the evolute pseudohastigerinid forms are closely related to P. micra but may represent a new species. Stable isotope data will also be analyzed in order to infer the depth ecologies of the various morphotypes.

Publication:

(1): Holladay, L. and Huber, B. In prep. (to be submitted to the Journal of Foraminiferal Research in fall 2001) Ontogenetic morphometric comparison of middle-late Eocene pseudohastigerinid planktonic foraminifera using X-radiograph image analysis.

(2): Results will be incorporated into the Atlas of Eocene Foraminifera (Huber, B. et al., in prep.).

Project Summary  |  Letter of Gratitude



Abby Knee

  • Gender: Female.
  • Ethnicity/Race: White.
  • Institution: University of Rhode Island.
  • Status: Junior.
  • Major: Marine Biology.

Career Goals: After earning a Ph.D., achieve a career focused on research in systematics and ecology of marine invertebrates.

Funding: Participation in the 2001 summer session of the Research Training Program was supported by the National Science Foundation Research Experience for Undergraduates program, Award: DBI-9820303.


Update - December 2002: Abby is in graduate school at Woods Hole Oceanographic Institution in the MIT/WHOI Joint Program, working toward a PhD in Biological Oceanography under the direction of Ken Halanych


Abigail J. Knee

Project Advisor: Dr. Kristian Fauchald, Department of Systematic Biology, Division of Worms.

Project Title: A cladistic analysis of the scale worm genus Halosydna.

Research Abstract: Historically, the annelid class Polychaeta, the bristleworms, has been wrought with misclassifications. The taxon has been presumed polyphyletic. Research focused on monophyly as a defining characteristic of clades has been problematic.

Within the polychaetes, the polynoid family of marine worms has particularly mystified taxonomists as this taxon resulted from the systematic removal of monophyletic groupings of scale worms from the former family encompassing them all.

The family Polynoidae consists of scale worms linked by the presence of scales and do not fit with the extracted taxa. Using cladistics and the premise that species represent descent with modification from a common ancestor, this research sought to determine the evolutionary relationships within the polynoid genus Halosydna Kinberg 1855 and in relation to its sister taxa.

The states of 88 morphological characteristics were described for eighteen species: eight species associated either historically or currently with Halosydna and ten species of closely related scale worms. Data was entered using the software DELTA and NDE. With this data, the computer software program PAUP produced 44 branching trees of relatedness revealing the most parsimonous paths of speciation based on the synapomorphies of the species.

Analysis of some cladograms in TREEVIEW revealed a grouping of Halosydna species within a monophlyletic clade, while those species removed to the genus Halosydnella fall out along various lines of descent.

Another notable pattern was the close grouping of Halosydna brevisetosa and H. leucohyba, proposed to be synonymous with each other. The resolution in the ingroup can now be further discussed so as to lead to evolutionary clarifications. In the course of the study, two undescribed species, H. augeneri and H. riojai, also were discovered in the NMNH worm collection, previously isolated by M. Pettibone but unpublished. The descriptions of these two species are to be submitted in the next year for publication in the Proceedings of the Biological Society of Washington.

Publication: Knee, A.J. and K. Fauchald. In Prep. A cladistic analysis of the genus Halosydna (Annelida: Polychaeta: Polynoidae), with a description of two new species: H. augeneri and H. riojai. Proceedings of the Biological Society of Washington.

Project Summary  |  Letter of Gratitude



Trevor Krabbenhoft

  • Gender: Male.
  • Ethnicity/Race: White.
  • Institution: North Dakota State University.
  • Status: Junior.
  • Major: Zoology.

Funding: Participation in the 2001 summer session of the Research Training Program was supported by the National Science Foundation Research Experience for Undergraduates program, Award: DBI-9820303.


Trevor J. Krabbenhoft

Project Title: Species-level systematics of the genus Symphurus (Cynoglossidae: Pleuconectiformes) of the western Pacific.

Research Abstract: The new species, described from 82 specimens collected during the Albatross
expeditions around the Philippine Archipelago in 1909, is characterized by a combination of characters including fin-ray counts, vertebral counts, and pigmentation features.

The new species has a predominant 1-2-2 pattern of interdigitation of dorsal pterygiophores and neuralspines and 14 caudal-fin rays.

The new species is most similar to S. woodmasoni, but differs drastically in its pigmentation.

Publication: Krabbenhoft, T. and T. Munroe. In prep. A new Cynoglossid Flatfish (Symphurus: Cynoglossidae: Pleuronectiformes) from Deep-waters of the Philippine Archipelago. Copeia.

Project Summary  |  Letter of Gratitude



Courtney Porreca

  • Gender: Female.
  • Ethnicity/Race: White.
  • Institution: University of New Mexico.
  • Status: Junior.
  • Major: Anthropology and Geology.

Career Goals: After obtaining a Ph.D., secure a position at a university teaching archaeology and conducting archaeological research.

Funding: Participation in the 2001 summer session of the Research Training Program was supported by the National Science Foundation Research Experience for Undergraduates program, Award: DBI-9820303.


Update - December 2002: Courtney attended the annual archaeology meetings (67th Annual Meeting of the Society for American Archaeology, Denver, 2002) and gave a paper on her RTP research.


Courtney A. Porreca

Project Advisor: Dr. J. Daniel
Rogers, Department of Anthropology.

Project Title: Paste Composition and Vessel
Function: A Case Study of Caribbean Ceramics.

Research Abstract: Archaeological ceramics provide a wealth of information about prehistoric cultures, including their technology. Prehistoric technology can also provide information on cultural traditions, lineages, and interactions, providing a broad picture of past cultures. Often the composition of the clay mixture used to make the pottery reflects technological choices made by the potter.

This study examines the relationship of this mix of ingredients and the intended use of the pottery from a period of almost 2,000 years on the Caribbean island of Nevis. To examine the clay mixture, a microscope was used to view cross sections of cut pottery sherds. The larger particles were identified and counted, and the form or shape of the vessel was used to infer its function. The time period or phase from which the vessels date was also a variable. The data was then analyzed to determine the relationship, if any, between the way the pottery was made and its intended use.

The results of this analysis show substantial continuity in the types and numbers of the inclusions across the different vessel functions as well as time periods. The same pattern emerged when the styles of surface treatments, for example painting or incising, were compared with the intended use of the pottery vessel. From this information, it can be concluded that the clay mixture of the ceramics from Nevis, as determined by counting the inclusions, is not directly related to either the intended function of the vessel or the decorative style. This is significant because it indicates a basic technological continuity on Nevis for almost 2,000 years despite theories of major cultural interaction due to migrations from mainland South America.

Future work that would provide more insight concerning this issue might include comparisons of the clay used to make the pottery to that of raw clay samples from the island, or comparing the overall Nevis ceramic collection to collections from other Caribbean islands.

Publication: Porreca, C. A. and J. D. Rogers. In prep. Paste Composition and Vessel Function: A Case Study of Caribbean Ceramics. Journal of American Archaeology.

Project Summary  |  Letter of Gratitude



Stacy Poulos

  • Gender: Female.
  • Ethnicity/Race: White.
  • Institution: University of Maryland.
  • Status: Non-graduating
    Senior.
  • Major: Anthropology and Art History.

Career Goals: After earning a Ph.D., attain a career as a research scientist at a museum focused on fieldwork in the Near East, with a concentration on archaeometric studies.

Funding: Participation in the 2001 summer session of the Research Training Program was supported by the National Science Foundation Research Experience for Undergraduates program, Award: DBI-9820303.


Update - December 2002: Stacy is currently in graduate school at the University of Pennsylvania studying Classics.

Anastasia L. Poulos

Project Advisor: Dr. Melinda Zeder, Curator of Zooarchaeology, Department of Anthropology.

Project Title: The Hunting of Gazella subgutturosa by Early Domesticators in the Fertile Crescent- A Comparison of Wild and Domestic Animal Remains in the Archaeological Record.

Research Abstract: The method of developing sex-specific age curves for domestic goat and sheep remains from archaeological sites has proved valid in identifying a kill-off profile that is representative of either domestication or hunting practices of people 10,000 years ago.

The application of this technique to the study of wild, hunted gazelle remains has further reinforced the potential for this kind of metric study, which is based on sexual dimorphism and fusion rates. Modern gazelle specimens were measured as a means of calibrating the degree of sexual dimorphism of the gazelle. Sexual dimorphism proved to be a dominant factor in the size of various bones. Regional variation was also strongly prevalent in the specimens; however, the ratio of male to female remained consistent and replicated the ratio of sexual dimorphism found in studies of modern goat and sheep.

This regional variation was found to be present in the archaeological record as well. Thousands of gazelle specimens were analyzed from archaeological sites throughout the highland and lowland regions of the Zagros in Iran û Ali Kosh, Asiab, Sarab, Guran, Palegawra, M'lefaat, and Jarmo. A comparison of the sex-specific survivorship curves derived from the archaeological data of hunted gazelle with that of domestic remains was a startling contrast. Domestic goats and sheep replicated a similar pattern of domestication practice to that of the previously studied site of domestication, Ganj Dareh, whereas the gazelle specimens gave strong evidence of a less specific selection in their hunt. The hunters focused more haphazardly, although there does seem to be a trend for the selection of bigger and older males, interspersed with female and young, which contrasts with the focused selection of young males for kill by the domesticators.

Project Summary  |  Letter of Gratitude



Yana Reid

  • Gender: Female.
  • Ethnicity/Race: Native American.
  • Institution: Haskell Indian Nations University.
  • Status: Junior.
  • Major: Biology with emphasis in Chemistry.

Career Goals: After obtaining a Ph.D., become a research scientist in the field of Ichthyology.

Funding: Participation in the 2001 summer session of the Research Training Program was supported by the National Science Foundation Research Experience for Undergraduates program, Award: DBI-9820303.


Update - December 2002: Yana spent the summer at OTS in Costa Rica and is now back in school. The results from her RTP research were recently submitted for publication.


Yana R. Reid

Project Advisor: Dr. Ron Heyer, Curator, Department of Systematic Biology, Vertebrate Zoology, DIvision of Amphibians and Reptiles.

Project Title: Analysis of advertisement calls in the genetically diverse frog taxon currently
known as Leptodactylus fuscus.

Research Abstract: The focus of this
project was to analyze the advertisement calls of the Leptodactylus fuscus.

Although morphologically very similar, previous electrophoretic analysis demonstrated the likelihood that this taxon contains more than one species. Using call analysis with Canary software along with the statistical application of SYSTAT, the call variation was quantified throughout the geographic range of L. fuscus.

Previous study of genetic differentiation in L. fuscus demonstrated that there was genetic partitioning among population units consistent with multiple species, rather than a single species, characterizing the unit currently understood as L. fuscus. Data were analyzed for 32 individual frogs from 24 localities throughout the distributional range of Leptodactylus fuscus. Data was taken on 10 calls for each frog (for those recordings with 10 or more calls). The data taken were call rate, call duration, carrier frequency, beginning frequency, ending frequency, frequency sweep, and temporal characterization of the waveform. Call rate, call duration, carrier frequency, ending frequency, and frequency sweep demonstrated a correlation with temperature and these parameters were standardized to 25C for further analysis.

Beginning frequency did not demonstrate a correlation with temperature. Multidimensional scaling analysis of our data indicated that frogs from the same locality showed the same kind of variation as frogs from different localities. The advertisement call data support the single species hypothesis in Leptodaactylus fuscus. Our study demonstrates that speciation in frogs is not always accompanied by differentiation of advertisement calls.

Project Summary  


  • Gender: Male.
  • Ethnicity/Race: Foreign, Hispanic.
Alexandre Cunha Ribeiro
Universidade de São Paulo
Ribeirão Preto, Brazil

Richard P. Vari, Ph.D.
Supervising Scientists
Department of Systematic Biology
Division of Fishes


"It was a great experience
working in my favorite
field, Phylogenetic
Biogeography, in one of
the best museums."


Phylogeny of the genus Oligosarcus Günther: A preliminary survey

The genus Oligosarcus comprises 16 species that occur in most of the major drainages of the south central portions of South America. Members of the genus are small to mid-sized fishes (about 40 to 200 mm) occurring mainly in streams. With the exception of O. schindleri, collected from a tributary of Río Charape which flows into the Rio Madeira of the Amazon basin, all Oligosarcus species occur outside the Rio Amazonas basin and no species of the genus have been collected north of 15º S (Menezes, 1988). Little ecological information on Oligosarcus is available although life history studies of a few species report that they range from omnivores to fish predators. Information on intrageneric relationships for Oligosarcus was effectively nonexistent and biogeographic hypotheses as a consequence were not grounded on phylogenetic information. Data from a series of osteological systems examined demonstrates that the genus is monophyletic, provides insight into likely sister groups to Oligosarcus, and partially resolves the interrelationships within the genus. The results of this study indicate that the fragmentation of the genus Oligosarcus into mostly allopatric species is congruent with a hypothesis of speciation associated with somewhat recent vicariant events as evidenced by the lack of sympatry between sister groups, a situation contrary to that in many other fish species in that region. The sister-group relationship between O. pintoi and O. planaltinae suggested by the results is consistent with the idea that the High-Paraná is a endemic area. However, the recognition of "lowland species" and "plateau species" groups as postulated by Menezes (1988) is not supported by the phylogeny, since some of the "plateau species" are more closely related to a subunit, but not all of the "lowland species."

This research was supported by the Alice Eve Kennington Endowment.

Project Summary  |  Letter of Gratitude



Julianne Rowehl

  • Gender: Female.
  • Ethnicity/Race: White.
  • Institution: Southampton College, Southampton, NY
  • Status: Graduated in 2001
  • Major: Marine Science with a Biology concentration

Career Goals: After obtaining a Ph.D., become a research scientist at a major university and study the ecology of estuarine or reef fish.

Funding: This student's participation in the summer 2001 Research Training Program was supported by a grant from the Smithsonian Women's Committee to provide an opportunity for international students and graduating seniors to participate in the program.


Rowehl, Julianne

Project Advisors: Dr. Carole Baldwin, Dr. Lynne Parenti, Dr. Victor Springer; Division of Fishes

Project Title: Identification of Galapagos Entomacrodus (Teleostei: Blennidae) with comments on relationships within the Entomacrodus "nigricans" species group

Research Abstract: Documenting biodiversity is important for understanding the complexity of ecosystems, as a foundation for future conservation decisions, and for understanding evolution in a historical context. Comparative morphological studies provide the basis for recognizing biological diversity. These studies involve careful comparisons of the anatomy of organisms, including superficial features and internal structures such as bones and cartilage. When a population of animals belonging to a known genus is discovered in a geographical area in which the genus has not been recorded previously, morphological comparisons with other known species of the genus are needed to identify the new population. In this study, fish of the tropical blenny genus Entomacrodus were collected from the Galapagos Islands in the eastern Pacific Ocean. This fish has never been reported from Galapagos. A comparative morphological study was performed, and the fish was identified as Entomacrodus chiostictus, a widespread species in the eastern Pacific. A study was then performed to examine relationships of 10 fish species within this genus. The main differences between the species in the genus are the coloration patterns, and detailed comparisons of numerous external and internal features in this study revealed few other differences. Color patterns are usually extremely important in fish because the females and males often recognize each other on the basis of these patterns. It has been hypothesized that Entomacrodus originated in the central Pacific Ocean and moved eastward to the eastern Pacific and Atlantic. Results of this study support a Pacific origin of the genus and suggest a close relationship among Atlantic members of the genus and E. chiostictus. Because the 10 species studied are so similar to one another anatomically, hypotheses of relationships within the group (based on morphology) are not well supported. Molecular data may provide more insight into the evolution of this fish genus.

Publication: None

Project Summary  |  Letter of Gratitude



Jeff Saarela

  • Gender: Male.
  • Ethnicity/Race: Foreign, White.
  • Institution: University of Manitoba, Winnipeg, Manitoba, Canada
  • Status: B. Sc. (Hons.) - 2000
  • Major: Botany

Career Goals: After obtaining a Ph.D., become a professor at a university and conduct research in plant systematics.

Funding: This student's participation in the summer 2001 Research Training Program was supported by a grant from the Smithsonian Women's Committee to provide an opportunity for international students and graduating seniors to participate in the program.


Saarela, Jeffery Michael

Project Advisors: Dr. Paul M. Peterson, Curator of Grasses, Department of Systematic Biology - Botany

Dr. Robert Soreng, Research Associate, Department of Systematic Biology - Botany;

Project Title: The taxonomy of Brachyelytrum (Poaceae).

Research Abstract: The Taxonomy of Brachyelytrum (Poaceae). Brachyelytrum, commonly known as long-awned wood grass, is a small native grass genus occurring in eastern North America and eastern Asia that grows in deciduous and mixed-coniferous forests. Workers in the past have recognized variation in the genus at the species, subspecies and varietal levels. Macromorphological and phytogeographical data support the recognition of three distinct species in Brachyelytrum. Brachyelytrum japonicum, distributed throughout southern Japan and eastern Korea, is characterized by narrow leaves [(0.5 0.7(0.8) mm], long first glumes [(0.2)(0.4-1.6)(2.0) mm] and long rachillas [((6.8)7.2-8.4(8.8) mm]. Brachyelytrum erectum is distributed throughout the eastern United States from northwestern Minnesota south to central Louisiana, east to northern Florida and north to northeastern New York. It tends to be a larger plant in many characters with respect to B. aristosum, and is characterized by longer hispid hairs on the lemma [(0.2)0.4-0.8(0.9) mm] and wide lemmas [(0.8)1.1-1.5(1.8) mm]. Brachyelytrum aristosum has a more northerly distribution, occurring in southern Canada from southern Ontario east to Newfoundland, the northern adjacent United States and south through the Appalachian mountains to northwestern North Carolina. It is characterized by short scaberulous hairs on the lemma [(0.06)0.08-0.14(0.2) mm] and narrow lemmas [(0.7)0.8-1.2(1.4) mm]. We include a key to the taxa, synonymies, representative specimens and descriptions.

Publication: Saarela, J.M., P.M. Peterson and R.J. Soreng. In Prep. (to be submitted fall 2001) The taxonomy of Brachyelytrum (Poaceae). Proposed for publication in the botanical journal Sida.

Project Summary  |  Letter of Gratitude



Shannon Schwaller

  • Gender: Male.
  • Ethnicity/Race: White.
  • Institution: St. Cloud State University.
  • Status: Junior.
  • Major: Anthropology.

Career Goals: Attain a Ph. D. in physical anthropology and then become a professor at a research university.

Funding: Participation in the 2001 summer session of the Research Training Program was supported by the National Science Foundation Research Experience for Undergraduates program, Award: DBI-9820303.


Shannon S. Schwaller

Project Title: Morphometric analysis of hand anatomy in five genera of New World primates.

Research Abstract: A morphometric analysis of hand and wrist bones in three genera of New World primates was conducted.

Specimens in sub-family Atelinae: Alouatta (species A.palliata and A. seniculus), Lagothrix (L. lagotricha) and Ateles (Ateles spp.) were compared with an out-group consisting of two genera of the sub-family Cebinae: Cebus (C. albifrons and C. capucinus) and Saimiri (S. sciureus). The closely related Atelinae have specialized hand morphology, such as elongated fingers and reduced thumbs. The Cebinae have a more generalized hand morphology. Comparing the Atelinae with an out group demonstrating less derived hand morphology tested the hypothesis that similar anatomical change in hand anatomy, such as finger length and thumb reduction, occurs in the same manner in closely related taxa.

The results from this study demonstrated that anatomical changes in proportions and growth allometry occur very differently in Ateles spp. relative to the other Atelinae species. Ateles spp. showed significantly longer metacarpal lengths relative to phalangeal lengths in the other Atelinae species. Also different in Ateles spp. was the reduction of thumb length through losses of both phalanges and in mid-shaft robustness through the loss of bony material in height and diameter. Ateles spp. demonstrated significantly different proportional and growth patterns of metacarpal length relative to phalangeal lengths from both species of Alouatta and the species L. lagotricha. Growth allometry in the long bones of the hand of Ateles differed significantly from the Atelinae. These results suggest that the genetic processes underlying the growth patterns have undergone significant changes among closely related species of the Atelinae sub-family.

Project Summary  |  Letter of Gratitude



Ainsley Seago

  • Gender: Female.
  • Ethnicity/Race: White.
  • Institution: Cornell University.
  • Status: Sophomore.
  • Major: Entomology.

Career Goals: After earning a PhD, achieve a career at a university teaching biology and/or entomology.

Funding: Participation in the 2001 summer session of the Research Training Program was supported by the National Science Foundation Research Experience for Undergraduates program, Award: DBI-9820303.


Ainsley E. Seago

Project Advisor: Dr. Steven Lingafelter, Department of Systematic Biology, Entomology.

Project Title: A New Species of Crepidodera (Coleoptera: Chrysomelidae).

Research Abstract: An unusual, seemingly polymorphic species of beetle was investigated and determined to represent an undescribed, cryptic species. Working with a generous series of suspiciously variable Crepidodera browni, we discovered a correlation between male genitalia shape and elytral color.

Through extensive dissection, examination, illustration, and analysis of these traits, we present a previously unknown species of flea beetle. We have name this minute, iridescent, coppery-bronze beetle Crepidodera praestigiator. C. praestigiator resides in the very backyard of the Smithsonian; the type specimens were found feeding on willow trees in the McKee-Beshers Wildlife Area in Montgomery County, Maryland.

108 beetles were dissected and their genitalia examined and illustrated. Crepidodera browni is described as a new species, found in Maryland and the Northeastern United States.

Publication: Seago, A. and S. Lingafelter. In Prep. A New Species of Crepidodera (Coleoptera: Chrysomelidae) from Maryland. Coleopterists Bulletin.

Project Summary  |  Letter of Gratitude



Dawn Stodden

  • Gender: Female.
  • Ethnicity/Race: White.
  • Institution: Western Montana College of the University of
    Montana.
  • Status: Non-graduating Senior.
  • Major: Environmental Science with emphasis in Geology.

Career Goals: After earning a Master's degree, pursue a career in field-based research
pertaining to conservation.

Funding: Participation in the 2001 summer session of the Research Training Program was supported by the National Science Foundation Research Experience for Undergraduates program, Award: DBI-9820303.


Dawn M. Stodden

Project Advisor: Dr. William Melson, Department of Mineral Sciences.

Project Title: The Assembly of North America: Using Garnet Zoning to Decipher the Geologic History of the Potomac Terrane.

Research Abstract: This study examined the
Sykesville Formation in the easternmost Potomac Terrain between Roosevelt Island and Chain Bridge in the Potomac River Gorge on the Virginia side. The Liberty Complex of the Potomac terrane is an accretionary prism (composed of the Sykesville Formation and the Morgan Run Formation) that was accreted on to Eastern North America probably over 400 million years ago by collision between a westward advancing magmatic arc and the accretionary prism.

This study of the Sykesville formation included structural mapping, precursory rock analysis, and chemical analyses of garnets and other minerals, all indicators of the geologic history of the rocks. The structural mapping the study area has shown a depositional trend that follows that of the surrounding terrain. In the Sykesville Formation, there are many quartz structures. These structures appear to be beds or veins. To decipher the protolith, thin section of quartz were analyzed using cathodoluminescence revealing relict sand grains preserved in the quartz thin section, giving evidence supporting that the quartz is depositional in nature. Garnets were also examined using electron microprobe analysis, a method in which the x-rays generated by a minute, 2-micron electron beam impinging on a mineral grain are analyzed for the elements that produced them. Chemical zoning in garnets acts as a record of the growth history of the garnet, as well as, the surrounding rocks. With the microprobe, several X-ray maps of some garnets in the Sykesville formation were constructed.

The research revealed that the Sykesville Formation, after deposition, has undergone one episode of metamorphism. Future analysis of the chemical composition of these garnets will unveil more about the pressures and temperatures at which the garnets were formed as well as the peak pressures and temperatures reached during the ancient collision, which added the Potomac Terrain to Eastern North America. With the data compiled in this study, more work to decipher the exact history of the Sykesville Formation will be possible.

Publication:

(1) Stodden, D. and W.G. Melson. In Prep. The Assembly of North America: Using Garnet Zoning to Decipher the Geologic History of the Potomac Terrane.

(2) Stodden, D. and W.G. Melson. 2002. Research Abstract, Southeastern Section, Geological Society of America, April 2002.

Project Summary  |  Letter of Gratitude



Katarina Topalov

  • Gender: Female.
  • Ethnicity/Race: Foreigh, White.
  • Institution: the University of Novi Sad, Yugoslavia.
  • Status: Junior.
  • Major: Plant Ecology and Animal Ecology

Career Goals: To obtain a Ph.D. in Botany and become a research scientist in the field of plant diversity

Funding: This student's participation in the summer 2001 Research Training Program was supported by a grant from the Smithsonian Women's Committee to provide an opportunity for international students and graduating seniors to participate in the program.


Katarina, Topalov

Project Advisor: Dr. Vicki Funk, Department of Systematic Biology, Botany.

Project title: Evaluation of Plant Diversity on the Guiana Shield

Research Abstract: The Guiana Shield is located in the northeastern part of South America and includes the countries Venezuela, Guyana, Surinam, French Giana, and the north part of Brazil. It hosts a unique assemblage of plants and animals that is extremely rich, but not well explored. Since this area is now threatened by resource extraction, it is important that we gain an understanding of its biodiversity so that decisions about its conservation can be made. In the past areas proposed for conservation have been difficult to evaluate because of incomplete data. There were few areas that have been collected well enough and the collections were lacking locality and other data partially or completely. In order to better understand the biodiversity the first goal of this project was to gather the data from a variety of sources and the second one was to evaluate the collecting bias such as time of year or intensity of collecting. For instance, one of the most commonly made mistakes in floristic analyses is the collecting and determining of only easily reached plant species (not far away from roads, tracks and rivers, etc.). However, even data like this can provide valuable information if analyzed properly, because they can be used as proxies in comparisons of two or more sites. The analyses included plant species checklist data from five areas: Puerto Ayacucho (Venezuela), Iwokrama Forest, Mabura Hill, Kaieteur Falls National Park (Guyana), Saül, Central French Guiana and Reserva Ducke, Brazil. The total of 5645 species records was standardized and imported into a database table. Finally, the ways of overcoming the collecting bias have been tested by using various kinds of statistical analyses in order to provide a more objective application of the already existing data and, therefore, more precise insight into the flora of northeastern South America. The gathered results were used to estimate plant diversity in western Venezuela (the Puerto Ayacucho area) and examine how this diversity compares to similar areas in French Guiana, Guyana and Brazil. The results of the study revealed that the flora of the Puerto Ayacucho area is the most different from those found on other five sites, but in order to bring more specific conclusions, more plant collecting expeditions have to be undertaken.

Publication:

(1): H. D. Clarke, K. Topalov, V.A. Funk and T. Hollowell. In prep. (to be submitted in fall 2001) Evaluation of Plant Diversity on the Guiana Shield.

(2): Proposed for publication in the Biodiversity and Conservation.

Project Summary  |  Letter of Gratitude



Sharon Wilson

  • Gender: Female.
  • Ethnicity/Race: White.
  • Institution: Middlebury College
  • Status: B.A. received May 2001
  • Major: Geology

Career Goals: After obtaining a Ph.D. in geology or planetary science, I plan to attain a research position at a university or governmental agency.

Funding: This student's participation in the 2001 summer session of the Research Training Program was supported by a grant from the Smithsonian Director's Office.

Wilson, Sharon

Project Advisor: Dr. Ed Vicenzi, Department of Mineral Sciences.

Project Title: Microscopy and microchemistry of low temperature aqueous alteration in the Lafayette meteorite

Research Abstract: Out of the 7500 meteorites that have been found on the surface of earth, only fifteen have been identified as Martian. The Lafayette meteorite is an achondrite that is almost certainly from Mars and was found in Lafayette, Indiana in 1931. The 800-gram stone has a crystallization age of 1.3 billion years and contains abundant hydrous secondary alteration veinlets that are 200-700 million years old. Although the age of the alteration is poorly constrained, it is clearly younger than the bulk rock. Since the underlying precept is that a Martian fluid created the low-temperature aqueous alteration, their physical and chemical characteristics provide crucial information in determining whether the Martian subsurface was once suitable to sustain microbial lifeforms. Hosted by iron-rich olivine and occurring along pyroxene grain boundaries, the veinlets possess a delicate microtexture consisting of silica-rich and iron-rich amorphous phases, smectite and carbonate. A detailed characterization of the textural and microchemical features of the alteration veinlets in Lafayette was undertaken. The olivine and pyroxene veinlets were analyzed by a sensitive X-ray microanalyzer (wavelength dispersive spectroscopy) to quantify the concentration of sulfur, sodium and chlorine in the secondary phases. The presence of sulfate is important regarding the search for life on Mars because it can be present in a form that allows microbial lifeforms to use it as an energy source. The presence of chloride in the alteration suggests that the water at or near the surface of Mars was slightly saline. The saline component would decrease the freezing temperature of liquid water, extending the conditions suitable for life on Mars.

Publication:

(1). Wilson, S.A. and E. Vicenzi. In Prep. (to be submitted December, 2001)

(2). Abstract proposed for publication at the Lunar and Planetary Science Conference, March 2002.

Project Summary  |  Letter of Gratitude


RESEARCH TRAINING PROGRAM
2001


STAFF

Director: Mary Sangrey
Assistant: Marita Davison


Program Summary
25 May 2002 - 3 August 2002

The 2001 Research Training Program opened the doors of the National Museum of Natural History to eighteen outstanding students selected from a highly competitive pool of applicants.

The students traveled from their home universities (both national and international) to take part in ten weeks of intense natural history research coupled with a thorough curriculum of lectures, tours, demonstrations, workshops, and field trips.

The primary focus of this summer's session was the realization of a research project, conducted under the personalized and skilled guidance of one or several research advisors, who served as mentors to each of the RTP students. Furthermore, the RTP curriculum covered the broad array of disciplines encompassed within the NMNH, offering the students a unique opportunity to experience and learn about subjects beyond their own personal interests.

The students journeyed through the extensive museum collections, viewed demonstrations of the museum's state-of-the-art technologies, and interacted with museum experts in the fields of Anthropology, Botany, Entomology, Invertebrate Zoology, Mineral Sciences, Paleobiology, and Vertebrate Zoology.

In summary, the 2001 Research Training Program offered the students an opportunity to build extensive research skills, interact with renowned scientists, and broaden their professional and personal horizons and served to fulfill the ultimate goal of contributing to the development of nascent natural history scientists.

STATS

Number of Lectures: 10
Number of Tours: 12
Number of Workshops: 7
Number of Demonstrations: 6
Number of Discussions: 8
Number of Field Trips: 3
Number of Socials: 3
Special Activities: 2
Number of hours dedicated to the research project: 255 hours/ 65%
Number of hours dedicated to other program events: 137 hours/ 35%


Research Activities and Finding

Research and Education Activities:

This is the third year of a 4-year Research Experiences for Undergraduates (REU) Site grant to support opportunities for undergraduate students to share in the research activities of Smithsonian scientists through participating in the Research Training Program at the Smithsonian Institution's National Museum of Natural History (NMNH) in Washington, D.C.

Supported by eleven Women's Committee grants (1983, 1991, 1993, 1994, 1995, 1996, 1997, 1998, 1999, 2000, and 2001), and totaling $170,200, 55 students have participated in the Research Training Program, including 4 in the '01 summer session.

The Research Training Program (RTP) is a ten-week, in-residence, summer program designed to train the next generation of scientists in collections-based research and natural history studies. The program incorporates research and study in Anthropology, Geology and Mineral Sciences, plus Systematic Biology, including Botany, Entomology, Invertebrate and Vertebrate Zoology. The program features REU student participants who are U.S. citizens, currently enrolled as an undergraduate student pursuing their first undergraduate degree. In addition, the RTP also complements REU positions by securing separate funding to support additional positions for undergraduate foreign students to join the site activities.

Students selected to participate in the '01 summer program received an award package of about $5,800, which included housing, transportation, research support budget, and stipend to cover living expenses.

The complete 2001 summer session of the Research Training Program hosted 18 undergraduate students, 12 of which were supported by this REU grant and 6 positions were supported by matching funds secured from other sources.

The ten-week program began 26 May 2001 and concluded 4 August 2001.

The complete RTP Class of '01 of 18 undergraduate students included 12 women and 4 individuals who identified themselves as members of ethnic or racial groups underrepresented in the sciences (minorities) including 2 who identified themselves as Native American and 2 who identified themselves as Hispanic. In addition, one participant indicated that they considered themselves physically challenged.

Of the 12 REU students included in the RTP Class of '01, 8 (67%) were women and 3 (25%) individuals identified themselves as members of ethnic or racial groups underrepresented in the sciences (minorities) including 2 Native Americans and 1 Hispanic. One REU participant indicated that they considered themselves physically challenged due to significant visual impairment.

The 2001 applicant pool totaled 219 applicants, of which 169 (63%) were women and 42 (16%) indicated in an optional questionnaire that they considered themselves an ethnic or racial minority. There was one applicant who indicated that they considered themselves physically challenged.

When considering students for placement, the RTP seeks to offer research opportunities to students who have not otherwise had the opportunity to connect with a scientific mentor or engage in research activities. The REU group of twelve '01 RTP participants included 1 Freshmen, 3 Sophomores, 6 Juniors, and 2 students who had completed 4 years of undergraduate study and therefore were classified at the Senior level but had at least one semester of undergraduate classes remaining after their participation in the '01 summer REU program.

The research activities of this REU site focused on undergraduate students conducting individual, hypothesis testing, research projects and participating in a curriculum of exploration through the biological, geological, and anthropological sciences including lectures, discussions, and workshops about research and careers in the sciences. Developing confidence and competence in the research process through direct interaction with a mentoring scientist continues to be the cornerstone of this program. This was accomplished by engaging student participants in the overall process of real life research from design to outcome including all the day-to-day steps in-between; offering scientifically significant research topics for students to pursue; and immersing them in a supportive scientific community of diverse researchers. All research project activities followed a common format. Students identified critical research questions, developed a specific hypothesis to test, designed and conducted original research, used the library, worked with the museum collections, gathered and interpreted data, and communicated their results to the scientific community through written and oral presentation. Students conducted their studies in-residence at the NMNH, focusing their activities on solving research questions by learning from the 124 million specimens that make up the Museum's national collections.

Complete abstracts of the research conducted by the students participating in the 2001 RTP is provided as part of the basic information reported for each student participant. In addition, research abstracts are posted on the web.


Research Findings

To emphasize the importance of communicating research findings, projects were designed to result in publication or presentation at a scientific meeting. Of the 18 research topics conducted in 2001, at least 16 publications or presentations are in various stages of reporting to the scientific community and anticipated to result in completion within the next 2-3 years. Some projects will require a longer time period to complete the analysis while other research topics represented preliminary investigations into new areas of science and therefore require further investigation before reporting.

Of the 12 research projects pursued by '01 REU students, 9 projects are anticipated to result in a total of 10 scientific works including 7 publications and 3 presentations at scientific meetings. These include:

Chacon, Y.J.*, and Dove, C.J., "Microscopic feather character analysis of the cranes, rails and allies (Order: Gruiformes)", Wilson Bulletin, vol., (), p. . In preparation, proposed for publication submission in 2003.

Rougvie, J.R., DiMaggio, E.N.*, Sorensen, S.S., "Polymetasomatic History of Ashflow Tuff and geochemical Evolution During Contact Metamorphism, Mammoth Crest, California", Geological Society of America 2001 Annual Meeting., vol. , (), p. . Accepted for presentation 2001.

Friedman, M.S.*, and G.D. Johnson, "A putative lampridiform from the Oligocene of Peru ", Journal of Vertebrate Paleontology, vol. , (), p. . In preparation, proposed for publication submission in 2002.

Holcomb, M.*, Pandolfi, J.M., Macintyre, I.G., and Budd, A.F., "Potential use of X-radiographs to distinguish members of the Montastraea annularis species complex.", Coral Reefs, vol. , (), p. . In preparation, proposed for publication submission in 2003.

Knee, A.J.* and K. Fauchald, "A cladistic analysis of the genus Halosydna (Annelida: Polychaeta: Polynoidae) with a description of two new species: H. augeneri and H. riojai.", Proceedings of the Biological Society of Washington, vol. , (), p. . In preparation, proposed for publication submission in 2003.

Krabbenhoft, T.J.* and Munroe T.A., "A New Cynoglossid Flatfish (Symphurus: Cynoglossidae: Pleuronectiformes) from Deep-waters of the Philippine Archipelago", Copeia, vol. , (), p. . In preparation, proposed for publication submission in 2003.

Porreca, C.A.* and J. Daniel Rogers, "Paste Composition and Vessel Function: A Case Study of Caribbean Ceramics", Journal of American Archaeology, vol. , (), p. . In preparation, proposed for publication submission in 2003.

Porreca, C.A.* and J. Daniel Rogers, "Paste Composition and Vessel Function: A Case Study of Caribbean Ceramics", Society for American Archaeology 2002 meetings. Poster and/or presentation, vol. , (), p. . In preparation. Proposed for presentation: 2002

Stodden, D.* and Melson W.G., "The Assembly of North America: Using Garnet Zoning to Decipher the Geologic History of the Potomac Terrane", Research Abstract, Southeastern Section, Geological Society of America, April 2002., vol. , (), p. . In preparation. Proposed for presentation: 2002.

Seago, Ainsley* and S. Lingafelter., "A New Species of Crepidodera (Coleoptera: Chrysomelidae: Alticinae) in Maryland.", The Coleopterists' Bulletin, vol. , (), p. . In preparation, proposed for publication submission in 2003.

* Designates REU student participant.


Training and Development

The Research Training Program aims to prepare undergraduate students for scientific careers by offering direct and personal first hand experience in the complete research process, focusing on a specific discipline, while also providing a curriculum of opportunities for exploration, and hands-on experience, across the expanse of natural history fields.

Activities conducted as part of the '01 summer REU program centered on attracting students to the field of natural history research, stimulating their interest in natural history research questions, preparing them for successful and productive careers, and highlighting the biocomplexities that provide linkages between disciplines.

The ambitious ten-week summer curriculum challenged participants to balance research time and time spent exploring other exciting opportunities available in natural history through program activities and Museum events. During the 2001 program students were in-session for 49 days (392 hours). An average of 32 days (255 hours) was reserved for dedicated project research (65%). The remaining time, approximately 17 days (137 hours), included about 51 different scheduled options such as lectures, tours, workshops, demonstrations, field trips, and social events. Options highlighted the National Museum of Natural History community and provided a unique peek into its' 124 million specimens and the research conducted by the 100+ in-residence scientific staff.

A complete archive of the 2001 Research Training Program is available on our web site at http://www.nmnh.si.edu/rtp/students/2001/01index.html The archive includes the schedule of activities, recruitment poster, and more.

The 2001 summer session of the Research Training Program hosted 18 students conducting 18 different natural history-based research projects under the mentoring guidance of 22 different research scientists. Of the 18 students participating in the program, 12 were supported by this NSF REU grant.

The '01 NSF REU students were:

Yolanda Jesusita Chacon*. 2001. New Mexico State University. Major: Biology. Academic Status: Sophomore. Research Project: Analysis of feather characters of cranes, rails, and allies (order Gruiformes). Research Advisor: Dr. Carla Dove, Department of Systematic Biology, Vertebrate Zoology, Birds. Gender: Female. Ethnicity/Race: Hispanic. Career Goals: After graduate school achieve a career in either Ornithology or Medicine. Web Posting: http://www.nmnh.si.edu/rtp/students/2001/students_2001_chacon.html

Erin Nicole DiMaggio*. 2001. University of Michigan. Major: Geology. Academic Status: Freshman. Research Project: Evidence for Feldspar Replacement in Altered Metamorphic Tuffs in the Duck Lake Volcanic Suite. Research Advisor: Dr. Sorena Sorensen, Department of Mineral Sciences. Gender: Female. Ethnicity/Race: White. Career Goals: After obtaining a Ph.D. in Geological Sciences, pursue a career as a volcanologist. Web Posting: http://www.nmnh.si.edu/rtp/students/2001/students_2001_dimaggio.html

Matthew Scott Friedman*. 2001. University of Rochester. Major: Biology and Geology. Academic Status: Junior. Research Project: Study of a putative lampridiform fish from the Oligocene of Peru. Research Advisors: Dr. James C. Tyler and Dr. G. David Johnson, Department of Systematic Biology, Vertebrate Zoology, Fishes. Gender: Male. Ethnicity/Race: White. Career Goals: After obtaining a Ph.D. in paleontology, achieve a career as a university professor conducting research on extinct and extant fishes. Web Posting: http://www.nmnh.si.edu/rtp/students/2001/students_2001_friedman.html

Michael Charles Holcomb*. 2001. University of Idaho. Major: Geology and Chemistry. Academic Status: Sophomore. Research Project: Use of X-radiographs for coral identification. Research Advisors: Dr. Ian MacIntyre and Dr. John Pandolfi, Department of Paleobiology. Gender: Male. Ethnicity/Race: Native American. Career Goals: After obtaining a Ph.D., obtain a position in an academic institution conducting research and courses on coral reef biology. Web Posting: http://www.nmnh.si.edu/rtp/students/2001/students_2001_holcomb.html

Abigail Jean Knee*. 2001. University of Rhode Island. Major: Marine Biology. Academic Status: Junior. Research Project: A cladistic analysis of the polychaete genus Halosydna. Research Advisor: Dr. Kristian Fauchald, Department of Systematic Biology, Invertebrate Zoology. Gender: Female. Ethnicity/Race: White. Career Goals: After earning a Ph.D., achieve a career focused on research in systematics and ecology of marine invertebrates. Web Posting: http://www.nmnh.si.edu/rtp/students/2001/students_2001_knee.html

Trevor James Krabbenhoft*. 2001. North Dakota State University. Major: Zoology. Academic Status: Junior. Research Project: A New Cynoglossid Flatfish (Symphurus: Cynoglossidae: Pleuronectiformes) from Deep-waters of the Philippine Archipelago. Research Advisor: Dr. Thomas Munroe, Department of Systematic Biology, Vertebrate Zoology, Fishes. Gender: Male. Ethnicity/Race: White. Career Goals: After earning a Ph.D., achieve a career focused on research in systematics of fishes. Web Posting: http://www.nmnh.si.edu/rtp/students/2001/students_2001_krabbenhoft.html

Courtney Ann Porreca*. 2001. University of New Mexico. Major: Anthropology and Geology. Academic Status: Junior. Research Project: Paste Composition and Vessel Function, A Case Study of Caribbean Ceramics. Research Advisor: Dr. Daniel Rogers, Department of Anthropology. Gender: Female. Ethnicity/Race: White. Career Goals: After obtaining a Ph.D., secure a position at a university teaching archaeology and conducting archaeological research. Web Posting: http://www.nmnh.si.edu/rtp/students/2001/students_2001_porreca.html

Anastasia Lee Poulos*. 2001. University of Maryland. Major: Anthropology and Art History. Academic Status: Non-graduating Senior. Research Project: The Hunting of Gazella subgutturosa by Early Domesticators in the Fertile Crescent, A Comparison of Wild and Domestic Animal Remains in the Archaeological Record. Research Advisor: Dr. Melinda Zeder, Department of Anthropology. Gender: Female. Ethnicity/Race: White. Career Goals: After earning a Ph.D., attain a career as a research scientist at a museum focused on fieldwork in the Near East, with a concentration on archaeometric studies. Web Posting: http://www.nmnh.si.edu/rtp/students/2001/students_2001_poulos.html

Yana Rae Reid*. 2001. Haskell Indian Nations University. Major: Biology. Academic Status: Junior. Research Project: Analysis of advertisement calls in the genetically diverse frog taxon currently known as Leptodactylus fuscus. Research Advisor: Dr. Ronald Heyer, Department of Systematic Biology, Vertebrate Zoology, Reptiles and Amphibians. Gender: Female. Ethnicity/Race: Native American. Career Goals: After obtaining a Ph.D., become a research scientist in the field of Ichthyology. Web Posting: http://www.nmnh.si.edu/rtp/students/2001/students_2001_reid.html

Shannon Scott Schwaller*. 2001. St. Cloud State University. Major: Anthropology. Academic Status: Junior. Research Project: A Morphometric Analysis of Hand Anatomy in Five Genera of New World Monkeys. Research Advisor: Dr. Richard Thorington, Department of Systematic Biology, Vertebrate Zoology , Mammals. Gender: Male. Ethnicity/Race: White. Career Goals: Attain a Ph. D. in physical anthropology and then become a professor at a research university. Web Posting: http://www.nmnh.si.edu/rtp/students/2001/students_2001_schwaller.html

Ainsley Ellen Seago*. 2001. Cornell University. Major: Entomology. Academic Status: Sophomore. Research Project: A New Species of Crepidodera (Coleoptera: Chrysomelidae). Research Advisor: Dr. Steven Lingafelter, Department of Systematic Biology, Entomology. Gender: Female. Ethnicity/Race: White. Career Goals: After earning a PhD, achieve a career at a university teaching biology and/or entomology. Web Posting: http://www.nmnh.si.edu/rtp/students/2001/students_2001_seago.html

Dawn Michele Stodden*. 2001. University of Montana, Western. Major: Geology. Academic Status: Non-graduating Senior. Research Project: The Assembly of North America: Using Garnet Zoning to Decipher the Geologic History of the Potomac Terrane. Research Advisor: Dr. William Melson, Department of Mineral Sciences. Gender: Female. Ethnicity/Race: White. Career Goals: After earning a Master's degree, pursue a career in field-based research pertaining to conservation. Web Posting: http://www.nmnh.si.edu/rtp/students/2001/students_2001_stodden.html


A profile of the four Women's Committee supported students follows.

Laura Holladay was a graduating senior from The University of Michigan where she earned an honors degree in the Geological Sciences with a senior thesis project on geomagnetism. For her RTP project, she worked with Dr. Brian Huber in the Department of Paleobiology studying the ontogenetic morphometrics of planktonic foraminifera of the genus Pseudohastigerina and an informally proposed new genus. The results of this work will be published in the Journal of Foraminiferal Research and incorporated into the forthcoming Atlas of Eocene Planktonic Foraminifera. An abstract of this work is published on the RTP web site at: http://www.nmnh.si.edu/rtp/students/2001/students_2001_holladay.html In the fall of 2001, she will begin graduate studies at the University of Michigan, and her funding will be provided by a National Science Foundation Graduate Research Fellowship. Her master's degree project, under the advisement of Dr. Kyger C. Lohmann, will be a
stable isotopic study of modern Adamussium colbecki, a scallop from the Antarctic. After receiving her M.S. at The University of Michigan, she plans to obtain a Ph.D. She will then pursue a career in teaching and research at an academic institution focusing on paleoclimatology and paleoceanography.

Julianne Rowehl was a graduating senior from Southampton College in Southampton, New York where she earned a degree in Marine Science with a concentration in Biology. Dr. Carole Baldwin, Dr. Lynne Parenti, and Dr. Victor Springer in the Division of Fishes collectively served as her team of research advisors. She studied the eastern Pacific blenniid fish Entomacrodus chiostictus and examined phylogenetic relationships in the Entomacrodus "nigricans" species group. An abstract of this work is published on the RTP web site at: http://www.nmnh.si.edu/rtp/students/2001/students_2001_rowehl.html After participating in the Research Training Program, Julianne plans on working for a year before applying to graduate schools where she anticipates studying fish ecology in estuarine or coral reef habitats. She would like to continue her education and receive a doctorate so she can serve as mentors to others while still conducting research.

Jeffery Michael Saarela graduated from the University of Manitoba, Winnipeg, Manitoba, Canada in 2000 with a B.Sc. (Hons.) in Botany. For his research project, he worked with Dr. Paul M. Peterson in the Department of Systematic Biology - Botany on the taxonomy of the grass genus Brachyelytrum (Poaceae). The results of this research will be published in the botanical journal Sida and is currently available on the RTP web site at: http://www.nmnh.si.edu/rtp/students/2001/students_2001_saarela.html. He plans to continue to collaborate with Peterson, working on a molecular characterization of the North American Brachyelytrum using DNA fingerprinting. After participating in the Research Training Program he began a M.Sc. program in September 2001 in the Department of Biological Sciences at the University of Alberta in Edmonton, Alberta, Canada. After graduate school he would like to obtain a university faculty position where he can teach and maintain an active research program in the field of plant systematics. Jeff says "The opportunity to work at the Smithsonian Institution with and among top-class researchers has been an incredible learning experience for me. The contacts I have made and the skills I have learned will certainly be an asset to my future career."

Katarina Topalov was a junior from the University of Novi Sad in Novi Sad, Yugoslavia where she has been studying Ecology and Environmental Studies. For her research project, she worked with Dr. Vicki Funk in the Department of Botany studying the biodiversity of the Guiana Shield. The results of their work will be published in the Biodiversity and Conservation Journal and an abstract is available now on the RTP web site at: http://www.nmnh.si.edu/rtp/students/2001/students_2001_topalov.html. After participating in the Research Training Program Katarina plans to finish her undergraduate degree at the University of Novi Sad and then proceed to a graduate program in botany. She would like to pursue a research career in the field of plant diversity.

Research investigations were conducted while students were in-residence at the National Museum of Natural History, 26 May 2001 through 4 August 2001.

An evaluation of the program, following a new method designed and implemented in 1999, indicates that all aspects of the 2001 summer program were successful in meeting Program goals and expectations. For a summary of the evaluation results visit our web page at: http://www.nmnh.si.edu/rtp/highlights/evaluations.html.


FINANCIAL SUMMARY

All 18 students participating in the 2001 summer session of the Research Training Program, including the 12 students supported by this NSF REU grant, each received the same award package totaling about $5,800 including a cash stipend paid directly to the student of $2,400. In addition to the stipend, the program provided participants housing and transportation to/from Washington, DC at no cost to the student participants. A research support budget was also granted to each student/mentor research team. The total award package proved adequate to cover all the student's expenses.

HOUSING: For housing, participants stayed at The Alexandria apartment complex in Alexandria, Virginia. A discounted group rate was negotiated between the Smithsonian and the complex to house the 18 students in four 4-person occupancy and one 2-person occupancy apartments for 10 weeks. The negotiated housing cost was $1,704 per person.

TRAVEL: All participants received a transportation allowance to accommodate their travel to/from Washington, DC. The average transportation costs for participants was $443.

RESEARCH SUPPORT: In addition to housing, stipend and transportation, each student/advisor team submitted a proposal outlining the research topic to be investigated and a supporting budget to cover the costs of the project. The average research support awarded was $516 per team. Through this funding option students were able to identify special activities to complement their research project and engage in additional enrichment opportunities.

Of particular note, 3 REU students were granted support to attend the Joint Meeting of Ichthyologists and Herpetologists, 1 REU student travel with their NMNH research advisor to their research project site for 2 weeks to collect additional specimens and record habitat information, and 1 REU student received funding to travel with their NMNH research advisor to another museum collection (Field Museum of Natural History) to study additional specimens included in their research.

OPERATIONAL COSTS: The cost to host the 2001 Research Training Program including costs to recruit student participants plus group expenses to implement the curriculum was $721 per student.


COST SHARING

The '01 summer Research Training Program was supported by three key grants including the NSF REU program, which contributed $76,200 (about 73%) to support, in part, 12 of the 18 undergraduate student participants.

The Institution secured the remaining $27,926, or 27% of the total costs to implement the 2001 summer session. The Smithsonian Women's Committee granted $18,800 (about 18% of the total) to support 4 students. An individual gift to the program by Alice Eve Kennington in 2000 also provided the foundation for a permanently endowed position in the summer session.

All staff salaries and facilities and instruction fees, laboratory and equipment fees, and general office supplies were contributed by the Institution at no cost to the program. The total secured to fully implement the 2001 Research Training Program including support for 18 undergraduate student participants was $104,126.


RECRUITMENT / SELECTION / EVALUATION PROCESS

The National Museum of Natural History, with no in-resident students, recruits all Program participants from outside the Institution. Broad recruitment using different media ensures that all undergraduate students, US and foreign, at large universities and small liberal arts and tribal colleges, as well as historically black colleges and universities, know about, and have the opportunity to apply for participation in the Program. For a summary of the recruitment efforts visit our web page at: http://www.nmnh.si.edu/rtp/highlights/recruitment.html and for specific demographic data about the 2001 effort review http://www.nmnh.si.edu/rtp/highlights/selecttables.html

Requests for information were received via e-mail, web, fax, phone, and general mail resulting in 219 proposals from students around the world, including 53 applicants representing 23 different countries, seeking to participate in the summer program (reference the complete data on the web at: http://www.nmnh.si.edu/rtp/highlights/international.html ).

To achieve this broad spectrum of students, representing some of the most promising young natural history researchers, recruitment posters were mailed to over 7,500 universities and institutions throughout the world. The mailing list included top ranking US and foreign universities as well as small colleges and target minority institutions. A concerted effort was made to reach out to potential participants from underrepresented groups through advertisement, participation in career fairs, and attending special conferences (e.g. Society for Advancement of Chicanos and Native Americans in Science (SACNAS)). At career fairs and conferences the Program hosted an exhibit booth, staffed by the Program Director, and participated in additional seminars and meetings to share information about opportunities for research, study and employment at the NMNH. Information was provided about the RTP specifically, and natural history in general. In addition, Program staff serve on a number of panels and belong to national organizations (e.g. Council on Undergraduate Research) to formally and informally promote the Program to students and professors at undergraduate universities.

To ensure that information and application materials are readily available to students, all information about, and application materials for, the Research Training Program are available on the World Wide Web (http://www.nmnh.si.edu/rtp/) including on-line fill-in forms and electronic submission capabilities. The RTP web site provides the option for students to prepare application materials on-line and submit them electronically.

For the 2001 Program 94% of all application materials were received on-line at this site, including the application form, cover letter, and letters of recommendation sent directly by references. To view the steady progression of on-line submissions see our web statistics at: http://www.nmnh.si.edu/rtp/highlights/webstats.html.

Students applying to the 2001 Program could also track their application on-line at our web site through the selection process and review research topics of students selected to participate. In addition, the site maintains a complete listing of Smithsonian advisors, their research specialties, former RTP students hosted, summaries and photos of their research topics sponsored, and abstracts of their presentations at scientific meetings and published papers. To view this information, visit our web page at http://rathbun.si.edu/rtp/application_procedure/advisor_list2.html

Of the 219 applicants to the 2001 program, 169 (63%) were women and 42 (16%) indicated in an optional questionnaire that they considered themselves an ethnic minority.There was one applicant who considered themselves physically challenged. Most applicants were US citizens, including at least 149 (68%). The majority of applicants to the '01 session were in their Junior year (31%). The 2001 applicant pool included 8 (4%) Freshman, 32 (15%) Sophomores, 68 (31%) Juniors, 25 (11%) non-graduating Seniors, 44 (20%) graduating Seniors, 16 (7%) students who had recently graduated, 6 (3%) graduate students, and 7 (9%) not listing their academic standing.

Most applicants to the '01 RTP indicated a preference for research topics in the biological sciences (39%) followed by the anthropological sciences (31%) and then the geological sciences (21%). Some, 20 (9%), expressed such diverse interests and project preferences that they could not be categorized into a specific discipline. The single most popular research topic was anthropology with 67 students selecting projects in this natural history discipline. The complete statistics are available at http://www.nmnh.si.edu/rtp/highlights/selecttables.html

The Research Training Program selection process is designed to identify students who will benefit the most from participating in the curriculum of the Program including those who are: motivated to participate in, and learn about, NMNH based research topics; interested in a career in the biological, geological or anthropological sciences (including research, teaching, or collections management); and equipped with sufficient education to understand the concepts and methodologies presented. The Program is also committed to recruiting and selecting women and minorities to serve as future role models in the natural history sciences.

To minimize bias, and to ensure that all applications are equally evaluated, the selection of participants follows an impartial seven step review process: 1) prescreening, 2) application scoring by scientific evaluators, 3) application pool normalization, 4) project advisor review, 5) finalist evaluation and scoring by scientific judges including participant nomination, 6) project advisor approval, and 7) applicant notification.

In summary, the 2001 selection process began with a pre-screening of all applicant files to eliminate incomplete, inappropriate, and non-competitive applications, at least one-third of the applicant pool. The pre-screening was conducted by two members from the NMNH scientific community who read and reviewed all applications, eliminating 54 (25%) and forwarding 165 (67%) as 'semi-finalists' to evaluators for further consideration. Sixteen members from the NMNH scientific community served as evaluators, assembled into eight review teams with two members per team. Each team reviewed 20 or 21 application folders representing a random set. Evaluators completed a 100 point score sheet for each application in their set. The review elements considered were: 1.) academic background, 2.) academic performance, 3.) science experience, 4.) motivation to participate in the RTP, 5.) career goals, 6.) career potential, 7.) letters of recommendation, and 8.) compensating characteristics / personal attributes (used to particularly credit unique or target applicants). Each application was scored by two different reviewers. Scores were normalized to eliminate scoring bias.

Based on the scores, the 60 top scoring applications were evaluated to ensure that there was adequate representation across research divisions and disciplines. To ensure a diverse group of undergraduate students, including representatives from different academic levels; ages and backgrounds; cultural, social, and ethnic groups; US states and territories; and foreign countries, some adjustments were made. The results yielded a finalist set of 50 applicants. After this initial ranking, potential research advisors were invited to review the entire applicant pool, including the finalist set, and advance into the final selection up to two applications uniquely suited to their research interest and projects.

A panel composed of seven scientists representing each of the NMNH research disciplines then reviewed all finalists. These seven judges nominated 18 student/advisor teams, including the 12 REU student/advisor teams, plus alternate teams. The judges also linked top applicants with appropriate mentors by matching mutual interests, prior experience, and stated preferences including the student's mentor selection. Mentors were then consulted about the judge's nominations and a few adjustments made to ensure a good student/mentor match. All applicants were notified of their final status 8 March 2001, as advertised. Invited participants and alternates were contacted personally by telephone to discuss placement and answer their particular concerns and questions. Three students declined participation, electing to join field research teams instead.

Most mentors initiated contact with their student immediately following selection to begin designing the research project.

The 2001 Program began Memorial Day weekend, 26 May 2001 and concluded ten weeks later, 4 August 2001.


EVALUATION PROCESS

After considerable research, the Research Training Program designed, developed, and implemented a new evaluation process in 1999 and continues to use this process to gauge the effectiveness of our program events and mentoring research advisors. The process provides a method and scoring system to evaluate success and quantitatively compare elements across years.

To monitor the design, content, and success of research topics each student/mentor team submits an initial proposal for the research project including a timetable for completion, budget and products anticipated. Proposals are reviewed by members from the NMNH scientific community and suggested adjustments implemented. To communicate research results, a formal seminar series is held with students presenting 15 minute illustrated, lectures. In addition, students prepare and present their research as part of a poster presentation and publish their research topics as a general abstract on the internet.


WOMEN AND MINORITY STUDENT PARTICIPANTS

The Research Training Program sought to improve minority representation in the applicant and participation pool by developing a partnership with the NSF Louis Stokes Alliances for Minority Participation program. Now in its fourth year, as a result of the LSAMP at the Smithsonian partnership, a more aggressive recruitment campaign was launched focusing on personal contact with LSAMP coordinators, multiple mailings to LSAMP centers plus their university partners, and internship positions for LSAMP scholars. We also hosted a very successful Winter Workshop in January '01 for students and site directors so as to help us develop an institutional plan for furthering the effectiveness of our outreach efforts into communities of underrepresented populations in the sciences.

The 2001 applicant pool of 219 included 42 (19%) minorities and out of the total 18 students who successfully competed to earn participation in the 2001 session of the Research Training Program, 4 were minorities (22%) - one being a student who participated in our January Workshop. In addition, the 2001 applicant pool included 58% women and women participants in the 2001 summer session of the Research Training Program represented 67%, or 12 of the total 18 students.


ADJUNCT ACTIVITIES

The curriculum designed for the 2001 Research Training Program was diverse. In addition to the research projects, events were selected to provide Program participants the opportunity to see all aspects of the NMNH. The complete schedule is available on our web site and can be seen at http://www.nmnh.si.edu/rtp/students/2001/schedule01.html and included 10 lectures, 12 behind-the-scenes tours, 7 workshops, 6 demonstrations, 8 discussions, 3 field trips, 3 socials and special activities.

Through a formal lecture series, Smithsonian researchers provided personal instruction in the concepts and techniques used in natural history research investigations. Training efforts focused on taking advantage of the wealth of natural history collections available at the Museum - 124 million specimens of plants, animals, anthropological artifacts, and geological samples. Through the curriculum events, staff shared insights into their collections-based investigations including details about how they manage their specific research programs and their personal contributions to science. In addition to lectures, weekly discussions and seminars were held to present topics in an open format. Topics covered included funding options for research and graduate school, the spectrum of job opportunities available in natural history disciplines, and personal insights about communicating research results including publishing and presenting. Rounding out the curriculum, workshops were offered on how to prepare, use, and maintain museum specimens. Behind-the-scene tours throughout the NMNH were hosted to broaden the participant's biological, geological, and anthropological perspective. The collection tours emphasized an appreciation of the diversity of the Earth and the role natural history museums play in studying and preserving that diversity. Weekend field trips to two of the Smithsonian's field research stations and one to a research site, Calvert Cliffs, accommodated a forum to discuss and demonstrate how natural history studies include both in the museum and field aspects and included collecting and observation techniques on-site. Laboratory demonstrations supported hands-on experience in various modern and traditional techniques covering many natural history sub-disciplines.

To teach the research process, under the guidance of a Smithsonian scientist, students participated in the design and execution of all phases of a scientific study including developing an explicit hypothesis, designing investigations to test this hypothesis, proposing a budget to accommodate the research, gathering data, making observations, analyzing results, preparing information for publication in a scientific journal and/or presentation at a scientific meeting. To insure an appropriate level of guidance, program staff monitored the progress of each research topic through a written hypothesis statement, mid-term evaluation, and final report and evaluation. Research projects focused on providing students training in how to communicate their research results, including producing a manuscript that will be accepted and published in a reputable, refereed, scientific journal and presenting the research topic to a scientific, and general, audience. To accommodate this training, at the conclusion of the summer program, students gave a 15-minute, illustrated, oral presentation in scientific meeting format on the progress of their research and prepared a final report in journal style. A poster session was also included to further illustrate the variety of mechanisms scientists utilize to communicate the research to a wide audience.

The curriculum of the program provided consistent training for all participants. Many research projects conducted during this award period focused on addressing the question 'What is a species?' and what types of characters and character states are used to distinguish between different species, thereby providing students training in traditional taxonomic techniques, phylogenetic analysis, theory and statistical methods of population comparisons. However, each research team included training elements unique to the sub-discipline or particular research topic.


Outreach Activities

Efforts are currently underway to seek additional avenues to expand the scope of science outreach to high school teachers, undergraduate and graduate students including expanding the suite of opportunities for them to participate in the research activities of Smithsonian scientists. Despite an apparent interest by Americans in the natural world, few undergraduate students are actively pursuing careers in the natural history sciences, such as systematic biology. Advertisement and recruitment for the Research Training Program served to not only attract a qualified pool of applicants, but also to heighten awareness of the research possibilities, and careers, in natural history disciplines.

By targeting efforts and special advertisements toward underrepresented groups such as African Americans, Hispanics, and Native Americans, we hope to encourage more minorities to achieve successful careers in the natural history sciences and thereby serve as role models for upcoming generations.

The Museum has also sought opportunities to serve as the venue for special science-based activities, thereby providing a forum to share our scientific expertise, rich collections, and facilities with larger audiences. For example, several NMNH research staff have secured grants from the National Science Foundation's Partnerships for Enhancing Expertise in Taxonomy (PEET) to directly participate in the education and hands-on training of a select group of students targeting careers aimed at taxonomic research of poorly understood organisms.

Recognizing that interest in natural history research does not necessarily begin at the undergraduate level but often stems from knowledge of, and positive experiences in the discipline during the high school and middle school years, the Museum has expanded its professional outreach to include younger audiences. Each year the Museum hosts a special behind-the-scenes tour for the 40 finalists in the Intel Science Talent Search, a national science competition for the nation's most promising science-talent high school students. Through an evening of tours and small group discussions with the finalists, NMNH scientific and collections staff share insights into their research activities, interesting research questions yet unanswered in the field of their expertise, and unique references available to researchers at the NMNH.

Products Produced

Dr. William Melson and undergraduate student Dawn Stodden have collected samples that will be valuable for future studies of this formation. There were six thin sections produced by this study team, as well as many field samples available for analysis. Also collected were structural data, chemical data, and X-ray maps. The data, field notes, and maps are key to future analysis of this formation.

From the research team of Dr. Steven Lingafelter and undergraduate student participant Ainsley Seago: 'A New Species of Crepidodera (Coleoptera: Chrysomelidae)' Series of Crepidodera were collected from Maryland and from Portal, Arizona. These specimens will be labeled appropriately and donated to the Smithsonian's entomology collection, where they will be freely accessible to the Museum's staff and visiting researchers.

From the research team of Dr. Daniel Rogers and undergraduate student Courtney Porreca: 'Paste Composition and Vessel Function: A Case Study of Caribbean Ceramics'. The following were collected: Data relevant to the sample and research carried out, in the form of aSystat database, and sample of cross sections of ceramics from Nevis. The data and the samples will become part of the complete Nevis material assemblage and possibly used in future research.

From the research team of Dr. Kristian Fauchald and undergraduate student Abigail Knee, 'A cladistic analysis of the polychaete genus Halosydna'. Created database of eight scored species of Halosydna. This file will be used by Dr. Kristian Fauchald in his network of scored specimens and may contribute to an interactive computerized key.

From the research team of Dr. James C. Tyler, Dr. G. David Johnson and undergraduate student Matthew Friedman, 'A putative lampridiform from the Oligocene of Peru'. The major products of the research include the preparation and tentative identification of a fossil skull that was unidentified and uncataloged prior to current examination. Furthermore, several illustrations of the skull and its more characteristic morphological features, as well as depictions of extant lampridiform fishes, were produced. The preparation and tentative identification of the fossil will be valuable to future researchers. Results from the research, including data and illustrations, will be made available to the scientific community upon the publication of our manuscript.

From the research team of Dr. Sorena S. Sorensen and undergraduate student Erin DiMaggio: 'Evidence for Feldspar Replacement in Altered Metamorphic Tuffs in the Duck Lake Volcanic Suite'. From this research two databases of the images were created. These images were gathered using Cathodoluminescence (CL) and SEM backscattered imaging as well as Microprobe X-ray maps. They are documented by their sample number and include all information collected. This database facilitates further analysis and comparison with remaining samples.

From the research team of Dr. Steven Lingafelter and undergraduate student participant Ainsley Seago*: "A New Species of Crepidodera (Coleoptera: Chrysomelidae)" Series of Crepidodera were collected from Maryland and from Portal, Arizona. These specimens will be labeled appropriately and donated to the Smithsonian's entomology collection, where they will be freely accessible to the Museum's staff and visiting researchers.

From the research team of Dr. William Melson and undergraduate student Dawn Stodden: 'The Assembly of North America: Using Garnet Zoning to Decipher the Geologic History of the Potomac Terrane'. Samples were collected from the research site that will be valuable for future studies of this formation. There were six thin sections produced by this study team, as well as many field samples available for analysis. Also collected were structural data, chemical data, and X-ray maps. The data, field notes, and maps are key to future analysis of this formation. As part of the US National Collections they are available to all researchers.

Contributions

Contributions within Discipline

The Smithsonian's National Museum of Natural History incorporates the largest and most comprehensive program of basic research in systematics and collections-based investigations in the world, embracing complex questions in the biological, geological and anthropological disciplines. The natural history sciences form an environment in which disciplines and sub-disciplines often cross-fertilize one another. Research topics pursued by students participating in the Research Training Program addressed key questions specific to individual fields, but often extended in an interdisciplinary fashion across areas of science. Students worked with their mentors employing a team approach to advance knowledge in a wide variety of disciplines. Studies applied both traditional and modern techniques.

Within the biological sciences there is a realization, shared by the public, of the worldwide crisis in rapid and uncontrolled loss of biodiversity. However, before greater advances can be made in conserving these diminishing resources, there is substantial 'basic' science that remains to be done to simply identify and catalogue this diversity. With 124 million specimens from which to learn, many RTP research topics in the biological sciences centered on questions of nomenclatural identity, taxonomic relationships, and evolutionary insights, thereby contributing to the descriptive knowledge of taxa and providing a foundation for more advanced and complex investigations in the future. Providing training in these basic studies is the major mechanism by which natural history museums, and programs such as the Research Training Program, contribute to the discipline of natural history.

Projects in the geological sciences also took advantage of the vast NMNH collections to expand classic studies, such as how fluids move through, and change, rocks during metamorphism.

The anthropological sciences cover a broad spectrum of topics from research directed toward developing a better understanding of physical collections to cultural issues and contexts.

Given the diversity of disciplines supported by this grant, and the multitude of research topics pursued, generalized summaries are reported with several individual examples. Many projects crossed disciplines in their scope and application.


Contributions to Other Disciplines

By definition natural history science incorporates a very broad spectrum of science. One of the goals of the Research Training Program is to provide undergraduate student participants an in-depth research experience in a particular discipline. As important, the program curriculum of lectures, discussion topics and tours through the natural history collections encourages students to consider linkages between disciplines plus extensions and applications beyond natural history science. In implementing such a curriculum, we hope to enhance the overall training of RTP student participants and to further their understanding of the biocomplexities in the sciences.

The complete curriculum for the '01 summer session can be viewed on our web site at: http://www.nmnh.si.edu/rtp/students/2001/schedule01.html


Ethics Component

New to the RTP suite of summer activities, the 2001 program included an ethics component focused on the conflict between freedom of access to museum specimen data and scientific confidentiality.

As part of the Smithsonian's National collections, more than 124 million specimens are available to the students for research and investigation. These collections have been acquired by the Museum for scientific study and represent historical specimens from the biological, geological, and anthropological disciplines. New technologies permit never-before-availability of these specimens, and their data, to scientists and general audiences from around the world. However, availability of specimen information, plus unrestricted access to scientific collections, poses important ethical decisions to be made by the new generation of natural history research scientists. Scientific collections can no longer be considered the independent property of the host institution and their scientific staff. New social opinions and governmental regulations draw into question: who has the authority to make these decisions?

To address these issues, as part of the summer schedule, a one-day workshop was held to actively engage students in thoughtful discussion and independent investigation. The complete workbook for this one-day workshop is posted on our web site at: http://www.nmnh.si.edu/rtp/students/2001/ethics01.html

Guided by an expert in the area, Dr. Lynn G. Clark, professor at Iowa State University, students evaluated two real-life case studies through a collective discussion, and then broke into small groups to comment on specific questions and apply parallel situations to their current REU project.

The discussion was lively and many issues were confronted that will
face the new generation of natural history scientists whose research
is dependent on the use of historic museum specimens plus acquisition
of new samples from worldwide localities. Following their discussion,
the students dispersed into the museum community to talk to
Smithsonian scientific staff about their experiences and gather
opinions. A debate was then held with students assigned to defend
either the pro (in favor) or con (against) of a proposition linked to
the issues of the workshop.

This format proved very effective in engaging students in forward
thinking about their important role as keepers of the new generation
of data, providing a format for them to openly discuss their thoughts
and learn the insights of established Smithsonian scientists, and then
to carefully look at both sides of the issue through the debate.

To illustrate the issues, students were posed with a real life
scientific dilemma guided by the researcher caught in the middle.
This approach proved exceptionally successful as students appreciated
that the topic was not a workbook exercise but a true and current
situation, still evolving, and with the outcome still pending.

The exercises dealt with a rare, endangered, and phylogenetically
critical species of plant known from only a few localities in Brazil.
Currently growing in a US research greenhouse from samples collected
years ago, and vouchered by several museum collections, specimens
representing the species are sought by horticulturists for commercial
trade and by researchers studying evolutionary lineages of the
monocots. Here called Species X, this plant is a potentially
profitable commodity and a critical piece in a yet unresolved
scientific puzzle. The species is a prime candidate for protection
under international endangered species regulations but is not yet
listed.

Key discussion points of the real-life ethics workshop included:

1. Scientific collections, such as those housed at the Smithsonian,
historically have been made available to those researchers with a
reasonable claim for data about the specimen. Recent technological
advances, including internet capabilities, now offers the opportunity
for scientific collections to be scanned and the complete specimen and
label data be visually available to anyone interested in viewing
collections. Should images of specimens like Species X be made
available to all audiences or should certain specimens or specimen
groups remain exclusive to scientific investigators only? What
criteria should be considered if limiting the availability of
information about certain specimens? Who will decide information
availability?

2. Scientific data, once published, are considered to be in the
public domain and data generated from federally funded research, such
as by Smithsonian research investigations, are regarded as public
property. Should the locality data of Species X be published, thereby
exposing it to potentially unscrupulous collectors including
overzealous biologists seeking specimens for research or
horticulturists anticipating profit from the sale of a rare and
endangered ornamental? What other natural history specimen locality
information poses potential problems if released into the easy-access
public domain?

3. If money to fund research and/or conservation efforts related to
Species X could be raised by selling plants grown in captivity, should
any of that money be returned to the government inwhich the species is
native? Is there a difference between exploiting the use of
endangered plant species for profit compared to animal species?

4. With such limited individuals of Species X - known from only three
populations and those located in high risk areas scheduled for
potential destruction - should museum collections be made to preserve
an adequate scientific record of Species X? How many collections
should be taken? Where should the collections be housed? Who will
determine the collecting limits and distribution of specimens?

5. Should a coordinated attempt be launched to preserve one or all of
the natural populations of Species X, even if publicity engenders
local hostility? Can international scientific interests supercede
local politics?

6. International and federal regulations governing species officially
listed as rare or endangered can inhibit research efforts by making it
extremely difficult to legally collect and import material. Will
formally listing Species X as being rare or endangered help or hinder
research efforts to understand and preserve the species?


Contributions to Education and Human Resources

The Research Training Program provides opportunities for undergraduate students to be directly involved in classic natural history studies and modern biological, geological, and anthropological research. Over the past twenty-one years, the program has developed into an internationally recognized opportunity for undergraduate students to gain background and experience in natural history research. By attracting promising students from diverse cultural and economic backgrounds who have a genuine interest in, and talent for, natural history-based research, and then providing these students the chance to explore and cultivate this interest by exposing them to the diverse possibilities available in the fields of biology, geology, and anthropology, we are contributing to the development of the human resource base of a globally-engaged next generation of scientists. In short, the Research Training Program provides unique insight into the life of a natural history research scientist for talented students who might otherwise never be exposed to research careers in the museum-based biological, geological, and anthropological sciences.

If Smithsonian research scientists are to affect the future of scientific investigation, and the development of human resources that support natural history studies, emphasis must not only be placed on communicating research results through written and oral presentation, but also on actively participating in the education, hands-on training, and inspiration of the next generation of natural historians. One of the best ways Smithsonian staff can contribute to the development of young professionals is by serving as mentors to students who come to the Museum to learn about research and career options. The Program offers to students, in the early stages of their professional training, the chance to be directly involved in natural history research. It also provides a forum for established Smithsonian scholars to serve as mentors sharing their experience, knowledge, and ideas with enthusiastic students who often possess the potential to become future leaders in the field, but lack the background and training to succeed.

Since the inception of the Research Training Program in 1980, and including the RTP Class of '01, 435 students have participated in the natural history research and systematic studies conducted by NMNH scientists and embarked upon the road to careers, many in these disciplines.

A complete directory of all RTP alumni, including their accomplishments since their summer at the Smithsonian, is now available on the web at http://www.nmnh.si.edu/rtp/alumni/directory.htm. This information is complemented by a photo gallery at http://www.nmnh.si.edu/rtp/alumni/alumniphotos.html and message center for posting new information at: http://www.nmnh.si.edu/rtp/alumni/messages.html

Maintaining statistics about the program and tracking students through their education and into careers following their participation in the Research Training Program has been an important element of the program. All program statistics are now maintained on our web site. For a directory of postings visit: http://www.nmnh.si.edu/rtp/highlights/statistics.html

Since the Program's founding, a total of 435 undergraduates have participated in the NMNH Research Training Program, including 252 (58%) women and 110 (25%) individuals identified as members of racial or ethnic groups underrepresented in the sciences (minorities). The complete summary is available at http://www.nmnh.si.edu/rtp/highlights/ed&car.html and http://www.nmnh.si.edu/rtp/highlights/partsum.html. Although we continue to seek new avenues to facilitate attracting and placing women and minorities in RTP positions, we believe we have identified many of the barriers that limits the access and success of these groups and therefore make every effort to avoid these.

To measure success, and evaluate the impact of the program, alumni follow-up and tracking has been an important component of the RTP and files show current data for 69% of participants. These statistics reveal that 78% of RTP alumni entered graduate programs in natural history research, including 56% of the minority alumni and 61% of the women with most pursuing degrees directly associated with, or complementary to, their experiences at the NMNH. Although many RTP alumni are still pursuing their education, of those not enrolled, 40% are employed in science careers and many, 38%, have returned to the NMNH for scientific investigation and to fill a variety of short and long-term positions. Through the use of our recently upgraded web site, that provides an easy and encouraging means for RTP alumni to keep us current on their progression, we hope to improve our alumni tracking and be better positioned to assess the impact our program has had on their education direction and career choices.


Contributions to Resources for Science and Technology

The majority of research projects conducted as part of the '01 summer session of the Research Training Program produced data resulting from the analysis of specimens from the Smithsonian's research collections. This data was then analyzed to draw conclusions based on the hypothesis being investigated. In some cases, additional museum specimens were collected, analysis samples prepared, and photographic or digital images produced.


Contributions Beyond Science and Engineering

Typically projects conducted as part of the Research Training Program do not offer contributions to the public welfare beyond science and engineering. None of the summer 2001 research teams reported implications of their study beyond the typical scope.

To date we have not implemented a Research Experiences for Teachers component but anticipate adding it to the '02 session.

Other collaborators

The Research Training Program is part of a larger initiative of the National Museum of Natural History to participate in the professional education and scientific training of the next generation of scientists in collections-based research and natural history studies including the biological, geological, and anthropological sciences. To support this initiative, the Smithsonian Institution provides a variety of direct and indirect support to facilitate the operation and implementation of the RTP and individual student positions.

The most significant in-house sources of financial support during this granting period have come from the Smithsonian Women's Committee ($18,800 - support for 4 students), and NMNH Office of the Director ($21,029 - program operation and implementation costs).

One position in the program was permanently endowed by Ms. Alice Eve
Kennington.

Contact with education and training organizations (e.g. Project Kaleidoscope (PKAL), Council on Undergraduate Research (CUR), Society for the Advancement of Chicanos and Native Americans (SACNAS)) including participation in their conferences and collaborations with their members, have helped shape the curriculum and direction of the Research Training Program.



ACTIVITIES AND FINDINGS


1. Describe the major research and education activities of the project.

From the research team of Dr. Carla Dove and undergraduate student Yolanda Chacon: 'Microscopic feather character analysis of the cranes, rails and allies (Order: Gruiformes).' The differences in microscopic characters of plumulaceous barbs have proven to be valuable in the identification of many different groups of birds. However, the role of the diagnostic characters in an evolutionary context is still unknown. This study aims to explore microscopic feather characters of the order Gruiformes by determining if the feather characters that are used in feather identification could be applied to phylogeny and evolution of birds. Eleven species of Gruiformes were selected from all representative families and one outgroup taxa from the Order Galliformes (Family: Phasianidae). Some of the microscopic plumulaceous feather characters examined in this study include: structures at node of the plumulaceous barbule base and tip, internode pigment at barbule base and tip, and symmetry of barbules. The key research activities included conducting a thorough literature search, preparation of microslides for identification, Scanning Electron Microscope (SEM) study, use of character matrix management, printing of trees and character matrix coding in MacClade 4.0 (Maddison, 2000) phylogenic software and the phylogenic trees were generated using phylogenic software PAUPÒ 4.0 (Swofford).


From the research team of Dr. Sorena S. Sorensen and undergraduate student Erin DiMaggio: 'Evidence for Feldspar Replacement in Altered Metamorphic Tuffs in the Duck Lake Volcanic Suite' Extensive studies on the alteration histories of volcanic rocks in eastern Sierra Nevada have been done by SI researchers. Although scientists know what rocks surround Duck Lake, until now no one had looked at the petrography of this group of metamorphic tuffs. The purpose of my project was to examine and analyze feldspar textures and mineral chemistry in order to determine the processes responsible for the alteration of these rocks. In addition, my results were compared with the chemical findings of the surrounding areas in order to determine how the geologic history of rocks in this area differed. In order to accomplish this three main instruments were used. First, and used most frequently, was cathodoluminescence (CL). I used CL to determine the growth and replacement history of feldspars in these rocks. With instrument assistance from my research advisor, Sorena Sorensen and Ed Vicenzi the CL proved to be a useful tool in identifying alteration patterns. SEM and Microprobe work was aided by Amelia Logan and James Rougvie. Using backscattered electron imaging and EDS analyses on the SEM I was able to qualitatively determine mineral composition in order to validate my CL observations and obtain images that would aid in navigating the microprobe. An electron microprobe was used to quantitatively determine mineral compositions and produce element X-ray composition maps. The X-ray maps were then used to analyze the location and distribution of major elements and were compared with CL and backscattered images. A scientific abstract was produced in collaboration with James Rougvie and Sorena Sorensen and submitted to the Geological Society of America for the 2001 annual meeting. Sorensen and Rougvie will continue work on the tuffs after the conclusion of this internship. In addition, my research will be incorporated in a larger scale study of the area and later published.


From the research team of Dr. James C. Tyler, Dr. G. David Johnson and undergraduate student Matthew Friedman: 'A putative lampridiform from the Oligocene of Peru'. A fossil fish skull (USNM 494403), tentatively identified by Bob Purdy and Dr. Jim Tyler as a luvarid, was examined in detail to more definitively determine its taxonomic status. The fossil was compared to skeletal, alcohol-preserved, and cleared and stained modern fish remains. Early in the process of comparing the specimen to luvarid material, it was apparent that its osteology was radically different from that of any described member of the suborder Luvaroidei, extant or extinct. The fossil was tentatively re-identified as a member of the order Lampridiformes based on an arrangement of the bones of the suspensorium and the opercular series that was strikingly similar to that of Lampris, a Recent lampridiform fish. In order to compare the specimen to members of the Lampridiformes, more of the fossil had to be exposed. The techniques utilized to reveal more anatomical details of the fossil skull included CT scanning, chemical preparation (acid treatment), and mechanical preparation. The most critical research activities involved mechanical removal of the matrix from potentially diagnostic areas of the skull that could prove useful in determining the taxonomic position of the fossil. These exposed areas were then compared to the equivalent regions in recognized lampridiform taxa. Modern lampridiform specimens for comparative work were taken from the collections of the National Museum of Natural History, the American Museum of Natural History, and the Scripps Institute of Oceanography. In addition to extensive preparation of the fossil and comparative work, the literature was searched thoroughly in order to provide data on modern and fossil lampridiforms. The osteology of the skull was described in detail, and several elements of the fossil were illustrated in order to highlight distinctive or potentially diagnostic morphological features. Further work, in particular more detailed CT scanning, needs to be performed in order to definitively place the fossil within the order Lampridiformes despite the very similar appearance of the fossil and some lower modern lampridiform taxa.


From the research team of Dr. John Pandolfi and Dr. Ian Macintyre and undergraduate student participant Michael Holcomb: "Use of X-radiographs for coral identification". Several characteristics were investigated for their potential in distinguishing members of the Montastraea annularis species complex in both living and fossil assemblages. Growth rates, corallite budding angles, curvature of growth bands, and band intensities were examined using X-radiographs and image analysis software. This data combined with locality data were used in an effort to assess the differences in members of the Montastraea annularis complex throughout their geographic range. The results show that the characteristics examined do show potential for being able to distinguish different species within the M. annularis complex, however a wider geographic area needs to be examined, genetic work needs to be done on the same specimens, and more detailed radiation measurements need to be obtained before the true potential of these characters can be assessed.

From the research team of Dr. Brian Huber and recent graduate Laura Holladay: 'Ontogenetic morphometric comparison of middle-late Eocene pseudohastigerinid planktonic foraminifera using X-radiograph image analysis'. In this study, biometric analysis of fossilized tests is used as the basis for classification of planktonic foraminiferal species. This project was designed to test the validity of an informally proposed new genus of planispiral foraminifera by comparing specimens referred to that genus with specimens assigned to a morphologically similar genus, Pseudohastigerina. As the latter genus is known to have a widespread global distribution, specimens for biometric analysis were chosen from an array of sites, including Mexico, Mississippi, an onshore borehole from New Jersey, and a deep-sea drill core from the tropical Pacific. One hundred twenty total specimens were chosen for x-radiography, as x-ray images allow observation and measurement of inner structures of the foraminiferal test. The x-rays were mounted on slides and then digitized using a video camera mounted onto a transmitted-light microscope. The digitized images were measured using biometric software. Measurements included the area of each chamber, the maximum test diameter, the total number of chambers present, the number of chambers in the initial and ultimate whorls, and the equation of the logarithmic spiral traced out by the series of planispiral chambers. Analysis of the biometric data supported distinction of two of the named species within the genus Pseudohastigerina but suggested that those specimens assigned to the species P. acutimarginata are probably synonymous with P. micra. Those specimens generally fell within the morphological variation of the species P. micra, and thus it is possible that they represent ecophenotypes of this wide-ranging species. A more evolute morphotype, however, may represent a new species within the genus Pseudohastigerina. As an extension of this project, stable isotopic analyses of these species are being conducted in order to test this hypothesis of ecophenotypic variation.


From the research team of Dr. Kristian Fauchald and undergraduate student Abigail Knee: 'A cladistic analysis of the polychaete genus Halosydna.' The family Polynoidae requires a major revision due to the inconsistencies of its previous taxonomic classification. This family consists of groups of scale worms left over from the removal of apparently uniform polychaete groups and is without any unifying features of its own. The polynoids cannot be shown as a monophyletic grouping and thus require further attention to delineate the contained species' relationships. For this study, ten specimens from the marine polynoid scale worm genus Halosydna were observed, and their characters were scored in order to evaluate the phylogenetic relationships between H. species, within the genus, and with eight outgroup taxa. Analysis should enable the researcher to revise the cladistic placement of H. spp. if necessary. Eighty-eight scored characters were entered into a data table in DELTA and NDE and run through the computer program PAUP to develop the most parsimonous cladograms. The characters included elytral patterns and ornamentation, prostomial features, chaetal ornamentation, and parapodial features. Thirty-five other characters were scored but not included in the PAUP analysis. Two undescribed species-H. augeneri and H. riojai-were discovered in the NMNH collection, and their descriptions are targeted for publication in the Proceedings of the Biological Society of Washington in 2002. Key research activities included conducting a thorough literature review, preparation of glycerin slides for observation of parapodia, observation of specimens under compound and dissecting microscopes, and data analysis. The results of this research support grouping Halosydna species within a monophlyletic clade, while those species removed to the genus Halosydnella fall out along various lines of descent. Another notable pattern was the close pairing of Halosydna brevisetosa and H. leucohyba, proposed to be synonymous with each other. The resolution in the ingroup can now be further discussed so as to lead to evolutionary clarifications. A PowerPoint slide show will be created and converted to photographic slides for an oral presentation of this study. The software PowerPoint will also be used to develop a poster for display at the RTP poster session.


From the research team of Dr. Thomas A. Munroe and undergraduate student participant Trevor J. Krabbenhoft: 'A New Cynoglossid Flatfish (Symphurus: Cynoglossidae: Pleuronectiformes) from Deep-waters of the Philippine Archipelago': The Indo-Pacific is home to over 20 nominal and several undescribed species of tonguefishes (Symphurus). Many of these are deep-water species that are known from only a few specimens. Our goal was to identify specimens that were originally thought to be Symphurus woodmasoni. In fact,we discovered that these were an undescribed species. We used traditional morphological characters in examining the specimens. We found that this species is characterized by a combination of a predominant 1-2-2 pattern of interdigitation of dorsal pterygiophores and neural spines; 14 caudal-fin rays, five hypurals, a black peritoneum, and numerous pigmentation characteristics.

From the Research Team of Dr. J. Daniel Rogers and undergraduate student Courtney Porreca: 'Paste Composition and Vessel Function: A Case Study of Caribbean Ceramics'. Ceramic assemblages provide a wealth of information about prehistoric cultures, including their technology. The ceramic assemblage from the Caribbean island of Nevis, which was collected through controlled excavation, is appropriate for a technological study because it offers a wide range of variation spanning a series of defined chronological periods. Of interest is the relationship between vessel function and variation in paste composition, specifically the inclusions, and whether new analytical methods can provide quantitative data useful for addressing comparative questions. Ceramicists generally accept that paste composition is central to the success of a vessel. In other words, the function of that vessel is dependent on the technology employed in the preparation of the paste. The method applied in this study involved counting individual inclusions, defined here as non-clay particles that occur naturally or were intentionally added to the clay, in a given area of a fresh cross-section. After identifying the common inclusions, three categories were created, light, dark and other, to facilitate the counting in an area of one square centimeter. An index value for quantity was calculated in an attempt to standardize the data. After performing basic statistical analyses, including correlations and t-tests, the results indicated substantial continuity across vessel form (from which function is inferred), and phase. A comparison of the exterior surface treatments with the paste composition revealed the same continuity. From this information, it can be concluded that the paste composition of ceramics from Nevis, as determined by counting the inclusions, is not directly related to the intended function of the vessel or the decorative style. This information is significant because it indicates technological continuity on Nevis for almost 2,000 years despite theories of major cultural interaction due to migrations from mainland South America. The method employed here is potentially very useful for future inter-island comparative studies of ceramic assemblages. Comparisons of the clay used in the sherds to that of raw clay samples from the island, as well as in increased sample size, would also be useful in examining the technology of production and its broader relationship to cultural interaction.


From the research team of Dr. Melinda Zeder and undergraduate student participant Anastasia Poulos: 'The Hunting of Gazella subgutturosa by Early Domesticators in the Fertile Crescent- A Comparison of Wild and Domestic Animal Remains in the Archaeological Record'. The method of developing sex-specific age curves for domestic goat and sheep remains from archaeological sites has proved valid in identifying a kill-off profile that is representative of domestication practices by people 10,000 years ago. The application of this technique to studying of wild, hunted gazelle remains has further reinforced the potential for this kind of metric study, based on sexual dimorphism and fusion rates. Thousands of gazelle specimens were analyzed from archaeological sites throughout the highland and lowland regions of the Zagros in Iran - Ali Kosh, Asiab, Sarab, Guran, Palegawra, M"lefaat, and Jarmo. A field expedition to the Field Museum of Natural History in Chicago provided much of the sample needed for this study. The collection of modern gazelle and archaeological remains at the Field Museum was extensive and was fully exploited by our research team. The measurements taken from long bones and other significant, sexually dimorphic bones (i.e., the astragalus) of the gazelle were all inputted into a database and manipulated for analysis through simple histograms and statistical programs.


From the research team of Dr. Carole Baldwin, Dr. Lynne Parenti, Dr. Victor Springer and undergraduate student Julianne Rowehl: 'Identification of Galapagos Entomacrodus (Teleostei: Blennidae) with comments on relationships within the Entomacrodus "nigricans" species group'. Documenting biodiversity is important for understanding the complexity of ecosystems, as a foundation for future conservation decisions, and for understanding evolution in a historical context. Comparative morphological studies provide the basis for recognizing biological diversity. During the 1998 Galapagos Imax Film Expedition, several specimens of the blenniid genus Entomacrodus were collected from shallow waters off Capo Douglas, Isla Fernandias. Although the specimens most closely resemble E. chiostictus, a widespread eastern Pacific species, this identification was questioned because the Galapagos Archipelago is well south of the known geographic range of E. chiostictus. Slight differences in pigmentation between the Galapagos specimens and E. chiostictus (noticeably the apparent absence of lip stripes) warranted morphological investigation to determine if the Galapagos population represents an undescribed species or if the geographic range of E. chiostictus includes Galapagos. No consistent differences in countable features such as fin rays, vertebrae, gill rakers, and pores were found between the Galapagos specimens and E. chiostictus. The greatest variation among eastern Pacific populations of Entomacrodus was observed in pigment patterns. No correlation between the various pigment patterns and any morphological feature was discovered, and all variation in pigmentation of the Galapagos specimens is accounted for in previous descriptions of E. chiostictus. Identification of the Galapagos specimens as E. chiostictus constitutes range extension for the species, which is now known to occur from the northern end of Isla Angel de la Guarda in the Gulf of Mexico to Playa Muerto, Panama including the offshore islands of Tres Marias, Revillagigedos, Clipperton, Cocos, and Isla Fernandias. Relationships among the 10 species of the Entomacrodus "nigricans" species group (sensu Springer), to which E. chiostictus belongs, were hypothesized based on cladistic analysis of external and osteological features. Central and western Pacific species are basal taxa in the phylogeny, supporting a previous hypothesis that the "nigricans" species group probably originated in the central Pacific Ocean and eventually moved eastward to the eastern Pacific and Atlantic. Atlantic and eastern Pacific Entomacrodus species form a monophyletic group. Because species of the "nigricans" group are extremely similar to one another anatomically, most nodes in the morphological - based phylogeny are not well supported. Future investigations involving molecular data may provide further insights into the evolution of Entomacrodus.


From the research team of Dr. Paul M. Peterson and undergraduate student Jeffery M. Saarela: 'The taxonomy of Brachyelytrum (Poaceae)'. Morphological variation in the genus Brachyelytrum has been variously treated at the species, subspecies and varietal level in the past. Nobody has conducted a detailed numerical analysis of the genus across its entire geographical range. In this study we performed a numerical analysis and a distributional study of Brachyelytrum from throughout its geographical range. The key research activities involved scoring 110 specimens of Brachyelytrum for 19 morphological characters, a scanning electron microscope study to search for micromorphological characters and to obtain micrographs for illustrative purposes, a literature search to obtain previous collection records, use of internet gazetteers to obtain latitude and longitude data, plotting specimen distributions in ArcView, and multivariate statistical analyses using discriminant analysis and principal components analysis to analyse the data set. In addition, we went on a four day collecting trip in Virginia, West Virginia, Maryland and Pennsylvania to observe living plants in the field, to collect voucher specimens, and to collect fresh material for future molecular studies.


From the Research team of DR. Richard Thorington and undergraduate student Shannon Schwaller: 'A morphometric analysis of hand anatomy in five genera of New World monkeys'. New World monkeys exploit a large variety of habitats and many species have evolved specialized hand adaptations. A morphometric analysis allowed for a detailed study of five different genera of New World monkeys. Woolly monkeys (genus Lagothrix), Howler monkeys (Alouatta) and Spider monkeys (Ateles), share many anatomical similarities, such as elongated fingers and small thumbs. This group with specialized hand anatomy was compared with a second set of monkeys, the Capuchin monkey (Cebus) and the Squirrel monkey (Saimiri), which have very little specialization. Measurements taken on twenty-nine monkeys from the specialized group were compared to the measurements of thirty-seven monkeys from the non-specialized group in order to test the hypothesis that similar anatomical changes in hand anatomy, such as the elongation of the fingers, occurs in the same way in closely related monkeys. Measurements of finger bone lengths and carpal bone articular surface sizes of the Atelinae genera were compared with the outgroup in order to determine morphological and development changes in Atelinae hand anatomy. Key research activities included a literature search to aide in research design, sorting and identifying hand bones for each genus examined, and data analysis using the Systat 9.0 statistical program. More comparative studies need to be done, especially together with the foot and with other New World monkeys, in order to understand better the genetic processes underlying the growth patterns observed in the Atelinae sub-family.


From the research team of Dr. Steven Lingafelter and undergraduate student participant Ainsley Seago: 'A New Species of Crepidodera (Coleoptera: Chrysomelidae)'. The most numerous and reliable species-specific differences within the members of a beetle genus are generally found in the genitalia; such differences serve a lock-and-key function to ensure intraspecific mating. By thoroughly examining the unique genitalia and elytral characteristics of two morphotypes within the species C. browni, this project distinguishes a new species of flea beetle, C. praestigiator. Key research activities included field observation and collection of Crepidodera, dissection of over 100 "C. browni" specimens, and microscope examination and camera lucida illustration of both male and female genitalia. Important characters of the male genitalia include degree of median lobe curvature, deflection of aedeagal apex, and shape of sac-eversion aperture. These traits are illustrated in shaded-graphite figures.


From the research team of Dr. William Melson and undergraduate student Dawn Stodden: 'The Assembly of North America: Using Garnet Zoning to Decipher the Geologic History of the Potomac Terrane'. Producing evolutionary histories of geologic formations provides valuable information about the past processes that affected the formations, as well as, giving evidence to the evolution of the continent in which the formation is present. There has been a lot of work done on the evolution of Eastern North America, as well as, the study area for this project, the Sykesville Formation. However, conclusive evidence as to a precise geologic evolution of the Sykesville Formation has not yet been presented. In order to do this study, samples were taken from the study area, which is from Roosevelt Island to Chain Bridge in the Potomac River Gorge on the Virginia side. Also recorded in the study area were structural data. By recording structural data present in the outcrops of our study area, the depositional history of this formation is becoming more understood. Also, by studying thin sections of quartz exposures from this study area with cathodoluminescence, evidence of the protolith of the quartz veins/beds has been found. Examinations of garnets found in the study area using electron microprobe analysis have given insight to the metamorphic history of the formation. Future calculations of chemical data collected will show more precisely the metamorphic history of the Sykesville Formation.


From the research team of Dr. Vicki Funk and undergraduate student Katarina Topalov: 'Evaluation of Plant Diversity on the Guiana Shield.' Various authorities have proposed that northeastern South America hosts a unique assemblage of plants and animals. This proposal has been difficult to evaluate in the past because of incomplete data. The research included plant species checklist data from five areas: Puerto Ayacucho (Venezuela), Iwokrama Forest, Mabura Hill, Kaieteur Falls National Park (Guyana), Saül, Central French Guiana and Reserva Ducke, Brazil. Finally, the ways of overcoming the collecting bias have been tested by using various kinds of statistical analyses in order to provide a more objective application of the already existing data and, therefore, more precise insight into the flora of northeastern South America. Research activities included standardization of a total of 5645 species records. All the data were imported into a database table and evaluated in several ways. Records for the Puerto Ayacucho area without latitude and longitude data but with a specific locality were georeferenced. The vegetation zones in the Puerto Ayacucho area were digitized on screen and the classifications added to the resulting attribute table. The results were used to estimate plant diversity in western Venezuela (the Puerto Ayacucho area) and examine how this diversity compares to similar areas in French Guiana, Guyana and Brazil. The results of the study did not correspond to a distance between any two sites or to commonly collected families. Therefore, they seem to have overcome collecting bias. The unique nature of all the sites shows that one of the sites cannot act as a surrogate for any of the others in terms of future conservation efforts.


From the research team of Dr. Ed Vicenzi and undergraduate student Sharon Wilson: 'Microscopy and microchemistry of low temperature aqueous alteration in the Lafayette meteorite'. The central aim of this research revolves around the characterization of low-temperature aqueous alteration veins and carbonate chemistry in the Lafayette Martian meteorite through detailed textural and microchemical analysis. The underlying precept is that a Martian fluid is responsible for the veins of aqueous alteration. Therefore, the physical and chemical characteristics of the secondary mineralization in meteorites provide crucial information to determine whether the environment beneath the surface of Mars was indeed within the "life window" as we have defined it with regard to terrestrial lifeforms. This research will help determine in further research if the alteration, typically hosted by iron-rich olivine, is the result of a single or multiple generations of fluid flow. The mineral chemistry of the clay associated with this fluid alteration will aid in identifying a terrestrial analogue environment, if one exists. Key research activities include a literature search and use of the polarizing microscope, environmental scanning electron microscope (ESEM) and the electron microprobe. From the microchemical results, I quantified the concentrations of sulfur, chlorine and sodium in the alteration veinlets, elements that enhance a life-sustaining environment. The presence of sulfur and chlorine may lead to the discovery of a new mineral. Lastly, the mineral composition of the carbonates is similar to carbonate alteration forming in young lava on the ocean floor at the Galápagos spreading center serving as a potential terrestrial analogue environment.


2. Describe the major findings resulting from these activities.

From the research team of Dr. Carla Dove and undergraduate student Yolanda Chacon: 'Microscopic feather character analysis of the cranes, rails and allies (Order: Gruiformes).' The analysis of the microscopic feather characters revealed that (1) feather characters do show some "natural groupings" among the taxa studied and may be aiding in the understanding of the phylogeny and evolution of birds (2) feather characters coupled with characters (i.e. morphology, DNA) may result in better resolved relationships; (3) a new research topic was discovered while conducting SEM analysis and involves determing whether certain feather characters are unique to the age of the feather, geography of the bird, or species. This study provided hands-on training in microslide preparation, scanning electron microscope analysis and microscopic feather character coding.


From the research team of Dr. Sorena S. Sorensen and undergraduate student Erin DiMaggio: 'Evidence for Feldspar Replacement in Altered Metamorphic Tuffs in the Duck Lake Volcanic Suite'. This research revealed that two phases of metasomatism altered the rocks from Duck Lake. The first was an early low-temperature alteration rich in potassium and sodium. The second alteration was fueled by a pluton driven calcic hydrothermal system. My results will help us understand how fluids move through rocks during metamorphism, and may eventually increase our understanding of ore deposit formation, tectonic history and the geochemical evolution of continental crust during magmatism.


From the research team of Dr. James C. Tyler, Dr. G. David Johnson and undergraduate student Matthew Friedman: 'A putative lampridiform from the Oligocene of Peru'. Though inconclusive, this research has established a striking similarity between the osteology of the fossil specimen and lower lampridiform taxa. Furthermore, the research produced a detailed description of the osteology of the skull as well as several illustrations of the more characteristic features of the fossil. If one tentatively accepts the lampridiform hypothesis, it is possible to place the fossil within the suborder Veliferoidei, the most basal suborder of the Lampridiformes, based on diagnostic morphological features. Further CT scan data, which will be available in the fall, should allow for the conclusive placement of the fossil within the order. Once this is accomplished, it seems likely that the fossil will be described as a new genus and species, given its age and unique suite morphological characters. This is a very significant find, as it is the only fossil lampridiform fish known from South America.


From the research team of Dr. John Pandolfi and Dr. Ian Macintyre and undergraduate student participant Michael Holcomb: "Use of X-radiographs for coral identification". This research revealed that several characteristics that are observable in X-radiographs do show statistically significant differences between different members of the Montastraea annularis species complex.


From the research team of Dr. Brian Huber and recent graduate Laura Holladay: 'Ontogenetic morphometric comparison of middle-late Eocene pseudohastigerinid planktonic foraminifera using X-radiograph image analysis'. The results of this biometric study suggest that an informally proposed new genus of planktonic foraminifera is actually composed of specimens that should be classified as the species Pseudohastigerina micra and a new species within the genus Pseudohastigerina.


From the research team of Dr. Kristian Fauchald and undergraduate student Abigail Knee: 'A cladistic analysis of the polychaete genus Halosydna'. This research revealed two new species to be described. The cladistic analysis support grouping Halosydna species within a monophlyletic clade, while those species removed to the genus Halosydnella fall out along various lines of descent. Another notable pattern was the close pairing of Halosydna brevisetosa and H. leucohyba, proposed to be synonymous with each other.


From the research team of Dr. Thomas A. Munroe and undergraduate student participant Trevor J. Krabbenhoft: 'A New Cynoglossid Flatfish (Symphurus: Cynoglossidae: Pleuronectiformes) from Deep-waters of the Philippine Archipelago': This research revealed that the specimens we examined were, in fact, an undescribed species. As such, we were able to formally describe the species.


From the Research Team of Dr. J. Daniel Rogers and undergraduate student Courtney Porreca: 'Paste Composition and Vessel Function: A Case Study of Caribbean Ceramics'. This research showed that the paste composition of ceramics from Nevis, as determined by counting the inclusions, is not directly related to the intended function of the vessel or to the decorative style. This information is useful because it indicates technological continuity on Nevis for almost 2,000 years despite theories of major cultural interaction due to migrations from mainland South America. The method employed here is potentially very useful for future inter-island comparative studies of ceramic assemblages.


From the research team of Dr. Melinda Zeder and undergraduate student participant Anastasia Poulos: 'The Hunting of Gazella subgutturosa by Early Domesticators in the Fertile Crescent- A Comparison of Wild and Domestic Animal Remains in the Archaeological Record'. The method of sex-specific age curves to denote domestic practices was applied to gazelle remains that were contemporaneous and contextually related to domesticated remains dating 10,000 years ago. This technique indicated that the kill-off pattern for hunting practices was very different from the kill-off of domesticated animals. With domesticates, young males were killed off, while females had a higher survival rate. Whereas, with the hunted remains, older, bigger animals were killed, with what appears to be a focus on males. The survivorship curves denote very different practices.


From the research team of Dr. Carole Baldwin, Dr. Lynne Parenti, Dr. Springer and undergraduate student Julianne Rowehl: 'Identification of Galapagos Entomacrodus (Teleostei: Blennidae) with comments on relationships within the Entomacrodus "nigricans" species group'. This research revealed that the Galapagos Entomacrodus constitutes a range extension for the species E. chiostictus. A comparative morphological study showed that the central and western Pacific species of the E. "nigricans" species group are basal taxa in the phylogeny. This supports a previous hypthesis that the group probably originated in the central Pacific Ocean and eventually moved eastward to the eastern Pacific and Atlantic. The Atlantic and eastern Pacific Entomacrodus species form a monophyletic group.


From the research team of Dr. Paul M. Peterson and undergraduate student Jeffery M. Saarela: 'The taxonomy of Brachyelytrum (Poaceae)'. Discriminant and principal components analysis of morphological data revealed three distinct groups correlated with geography. We recognize three distinct species: B. japonicum from Asia, and B. erectum and B. aristosum from North America. Brachyelytrum japonicum is characterized by its narrow leaves, long first glumes and long rachillas. Brachyelytrum erectum is characterized by long hairs on the lemma, wider lemmas and wider on average leaves and is distributed throughout the eastern United States from the Canada-U.S. border south to Florida. Brachyelytrum aristosum is characterized by short hairs on the lemma, narrower lemmas, and narrower on average leaves and has a more northerly distribution, occurring in southern Canada, northern United States and south through the Appalachian mountains.


From the Research team of DR. Richard Thorington and undergraduate student Shannon Schwaller: 'A morphometric analysis of hand anatomy in five genera of New World monkeys'. This research revealed that the genus Ateles demonstrated very different anatomical changes in proportions and growth patterns compared to the other Atelinae genera. Ateles showed significantly longer metacarpal lengths relative to phalangeal lengths. Ateles also demonstrated a very different growth allometry in finger development compared to the Atelinae and outgroup species.


From the research team of Dr. Steven Lingafelter and undergraduate student participant Ainsley Seago: 'A New Species of Crepidodera (Coleoptera: Chrysomelidae)'. Results of this study confirmed the hypothesis that a new species of Crepidodera, previously grouped as a morphotype of C. browni, occurs in Maryland.


From the research team of Dr. William Melson and undergraduate student Dawn Stodden: 'The Assembly of North America: Using Garnet Zoning to Decipher the Geologic History of the Potomac Terrane'. This research has shown that the trend of the rocks present in the Sykesville Formation follows the trend of the surrounding terrain. Also discovered were relict sand grains in quartz beds that were previously described as veins. Finally, the chemical zoning in the garnets present in this formation have shown that the formation experienced only one metamorphic episode.


From the research team of Dr. Vicki Funk and undergraduate student Katarina Topalov: 'Evaluation of Plant Diversity on the Guiana Shield.' The results of the study did not correspond to a distance between any two sites or to commonly collected families. Therefore, they seem to have overcome collecting bias. The unique nature of all the sites showed that one of the sites cannot act as a surrogate for any of the others in terms of future conservation efforts.


From the research team of Dr. Ed Vicenzi and undergraduate student Sharon Wilson: 'Microscopy and microchemistry of low temperature aqueous alteration in the Lafayette meteorite.' Through this research I characterized the alteration veinlets in Lafayette and quantified the concentrations of sulfur, chlorine and sodium in the carbonate, clay and amorphous phases which may lead to the discovery of a new mineral.


3. Describe the opportunities for training and development provided by your project.


From the research team of Dr. Carla Dove and undergraduate student Yolanda Chacon: 'Microscopic feather character analysis of the cranes, rails and allies (Order: Gruiformes).'This study provided specialized training in conducting a thorough literature search, preparation of microslides for identification, Scanning Electron Microscope (SEM) study, use of character matrix management, printing of trees and character matrix coding in MacClade 4.0 (Maddison, 2000) phylogenic software and the phylogenetic trees generated using phylogenetic software PAUPÒ 4.0 (Swofford).


From the research team of Dr. Sorena S. Sorensen and undergraduate student Erin DiMaggio: 'Evidence for Feldspar Replacement in Altered Metamorphic Tuffs in the Duck Lake Volcanic Suite'. This research provided training in Cathodoluminescence (CL) imaging, SEM backscattered electron imaging and EDS analysis, and Electron Microprobe analysis with distribution X-ray maps. This training will be instrumental in my future career as a research scientist.


From the research team of Dr. James C. Tyler, Dr. G. David Johnson and undergraduate student Matthew Friedman: 'A putative lampridiform from the Oligocene of Peru'. This research provided experience useful to one pursuing a career in paleoichthyology. Working directly with Dr. Tyler and Dr. Johnson provided valuable knowledge of fish osteology, while Dr. Tyler was an excellent source of instruction regarding scientific illustration. Being able to present renderings of one's objects of study is clearly an important skill in many scientific fields, and paleoichthyology is no exception. Basic instruction on mechanical and chemical preparation techniques for fossil remains as provided by preparator Fred Grady was instrumental in completion of this project, and the training received is valuable in many paleontological applications. Introduction to the use of CT scanning technology was another byproduct of this research, and will be useful in future studies of fossils.


From the research team of Dr. John Pandolfi and Dr. Ian Macintyre and undergraduate student participant Michael Holcomb: "Use of X-radiographs for coral identification". This project provided an opportunity to become more familiar with both image and statistical analysis software.


From the research team of Dr. Brian Huber and recent graduate Laura Holladay: 'Ontogenetic morphometric comparison of middle-late Eocene pseudohastigerinid planktonic foraminifera using X-radiograph image analysis'. This research provided specialized training in the preparation of microfossil samples, acquisition of x-radiograph images (including x-ray film developing), preparation of slides, use of computerized video capture and digitizing tablets for digital image production, use of biometric software, and data analysis. These techniques were learned in the context of species-level identification of microfossils.


From the research team of Dr. Kristian Fauchald and undergraduate student Abigail Knee: 'A cladistic analysis of the polychaete genus Halosydna'. This research provided training in conducting literature reviews, morphological polychaete systematics, character scoring, data analysis, cladogram interpretation, and preparation of the description of two new scale worm species for publication.


From the research team of Dr. Thomas A. Munroe and undergraduate student participant Trevor J. Krabbenhoft: 'A New Cynoglossid Flatfish (Symphurus: Cynoglossidae: Pleuronectiformes) from Deep-waters of the Philippine Archipelago'. This research provided training in morphological fish systematics,observational techniques literature evaluation and review, data analysis, and preparation of the scientific description of a new species of fish for publication.


From the Research Team of Dr. J. Daniel Rogers and undergraduate student Courtney Porreca: 'Paste Composition and Vessel Function: A Case Study of Caribbean Ceramics'. This research provided training in ceramic analysis as well as the development and execution of a research project. Minimal training with Cathodoluminesence, the Polarizing Microscope and the Scanning Electron Microscope was also received. The results contribute to the existing knowledge of the prehistory of the Caribbean.


From the research team of Dr. Melinda Zeder and undergraduate student participant Anastasia Poulos: 'The Hunting of Gazella subgutturosa by Early Domesticators in the Fertile Crescent- A Comparison of Wild and Domestic Animal Remains in the Archaeological Record'. This project provided specialized training in recognizing goat, sheep, and gazelle species, as well as discerning fusion rates and tooth wear patterns, all of which are important in archaeological analysis of a site.


From the research team of Dr. Carole Baldwin, Dr. Lynne Parenti, Dr. Springer and undergraduate student Julianne Rowehl: 'Identification of Galapagos Entomacrodus (Teleostei: Blennidae) with comments on relationships within the Entomacrodus "nigricans" species group'. This research provided specialized training in the preparation of cleared and stained specimens, fish osteology, character development and scoring, and use of PAUP 3.0 and MacClade 3.0.


From the research team of Dr. Paul M. Peterson and undergraduate student Jeffery M. Saarela: 'The taxonomy of Brachyelytrum (Poaceae)'. This research provided training in morphological plant systematics, including character development and scoring, preparation of distribution maps, scanning electron microscopy, data analysis using multivariate statistics, and field collecting techniques.


From the Research team of DR. Richard Thorington and undergraduate student Shannon Schwaller: 'A morphometric analysis of hand anatomy in five genera of New World monkeys'. This research provided training in primate anatomy, design of a morphometric study, and data analysis using computer statistical programs.


From the research team of Dr. Steven Lingafelter and undergraduate student participant Ainsley Seago: 'A New Species of Crepidodera (Coleoptera: Chrysomelidae)'. This research provided training in entomological field identification, collection, microdissection, and illustration techniques, as well as preparation of a scientific paper for publication.


From the research team of Dr. William Melson and undergraduate student Dawn Stodden: 'The Assembly of North America: 'Using Garnet Zoning to Decipher the Geologic History of the Potomac Terrane'. This research provided training in structural measurements and analysis, cathodoluminescence imaging and interpretation, SEM imaging and analysis, electron microprobe use and analysis, and analysis of chemically zoned minerals. Training in skills needed to present research for publication was also part of the project.


From the research team of Dr. Vicki Funk and undergraduate student Katarina Topalov: 'Evaluation of Plant Diversity on the Guiana Shield.' This research provided training in data basing and analyzing, the GIS course in georeferencing, using the GPS as a valuable tool for botanical studies, plant collecting and preservation, and preparation of the scientific paper for publication and audio and visual presentation.


From the research team of Dr. Ed Vicenzi and undergraduate student Sharon Wilson: 'Microscopy and microchemistry of low temperature aqueous alteration in the Lafayette meteorite.' This research provided experience using the polarizing microscope, training in ESEM imaging and analysis and electron microprobe use and analysis. Software programs such as analySIS and SigmaPlot were also utilized.


4. Describe outreach activities your project has undertaken.


From the research team of Dr. Sorena S. Sorensen and undergraduate student Erin DiMaggio: 'Evidence for Feldspar Replacement in Altered Metamorphic Tuffs in the Duck Lake Volcanic Suite'. The results of this research produced an abstract that was submitted to the Geological Society of America for the annual meeting in Boston, Massachusetts, November 2001 and will be presented as a talk. The abstract and the talk are available to the public, a sharing of ideas that will serve as a stimulant for further support of research.


From the research team of Dr. James C. Tyler, Dr. G. David Johnson and undergraduate student Matthew Friedman: 'A putative lampridiform from the Oligocene of Peru'. The results of the study will be presented at the University of Rochester to non-scientist members of the student body during a colloquium on undergraduate research. Furthermore, similar presentations are planned for science classes at the author's high school to encourage interest in the natural sciences among younger students.


From the research team of Dr. John Pandolfi and Dr. Ian Macintyre and undergraduate student participant Michael Holcomb: "Use of X-radiographs for coral identification". A presentation covering the project as well as the RTP program in general will be given to a geology class at the University of Idaho.

From the research team of Dr. Kristian Fauchald and undergraduate student Abigail Knee: 'A cladistic analysis of the polychaete genus Halosydna'. Gave three tours of research project and collection to other Smithsonian science interns and visitors.


From the Research Team of Dr. J. Daniel Rogers and undergraduate student Courtney Porreca: 'Paste Composition and Vessel Function: A Case Study of Caribbean Ceramics'. This research resulted in a paper proposed for publication and a poster. A formal presentation was also given to an audience of research scientists.


From the research team of Dr. Paul M. Peterson and undergraduate student Jeffery M. Saarela: 'The taxonomy of Brachyelytrum erectum Schreb. (Beauv.) (Poaceae)'. An oral summary of this research project was provided to interns from the Smithsonian Environmental Research Centre (SERC) and the Conservation Resource Centre (CRC)) during their tour of the National Museum of Natural History.


From the research team of Dr. Steven Lingafelter and undergraduate student participant Ainsley Seago: 'A New Species of Crepidodera (Coleoptera: Chrysomelidae)'. During field collection, some of our entomological goals were informally described to inquisitive local nonscientists.


From the research team of Dr. Ed Vicenzi and undergraduate student Sharon Wilson: 'Microscopy and microchemistry of low temperature aqueous alteration in the Lafayette meteorite.' The results from this research will be published in the form of an abstract and poster at the Lunar and Planetary Science Conference in March 2002. The abstract and poster are available to the public and may inspire new ideas for future research.



CONTRIBUTIONS

1. Contributions to the principal disciplines(s) of the project


From the research team of Dr. Carla Dove and undergraduate student Yolanda Chacon: 'Microscopic feather character analysis of the cranes, rails and allies (Order: Gruiformes).' This study provides a foundation for applying feather characters in phylogeny and evolution of birds. Although more research is needed on the feather characters in this group of birds, this preliminary study provides some evidence that feather characters produce trees that cluster into "natural groupings" in parsimony analysis and supports the idea that these characters can be useful for identification and phylogenetic analysis.


From the research team of Dr. Sorena S. Sorensen and undergraduate student Erin DiMaggio: 'Evidence for Feldspar Replacement in Altered Metamorphic Tuffs in the Duck Lake Volcanic Suite'. This research shows how fluids move through and change rocks during metamorphism. It shows both the textural and chemical alterations that occur in both low and high temperature environments including the impacts of hydrothermal systems.


From the research team of Dr. James C. Tyler, Dr. G. David Johnson and undergraduate student Matthew Friedman: 'A putative lampridiform from the Oligocene of Peru'. This research contributes data to the scant fossil record of fishes of the order Lampridiformes. Since the modern members of the order are so morphologically disparate, the evolutionary history of the group is of particular interest. Furthermore, the specimen is the only known fossil lampridiform from South America.


From the research team of Dr. John Pandolfi and Dr. Ian Macintyre and undergraduate student participant Michael Holcomb: "Use of X-radiographs for coral identification". The addition of more characteristics distinguishing members of the Montastraea annularis species complex will be of value in the identification of specimens that are currently unidentifiable due to insufficient distinguishing characteristics. This is useful in understanding paleoecology and the evolutionary history of the M. annularis complex.


From the research team of Dr. Brian Huber and recent graduate Laura Holladay: 'Ontogenetic morphometric comparison of middle-late Eocene pseudohastigerinid planktonic foraminifera using X-radiograph image analysis'. This research provides an example of a carefully structured biometric analysis for assessing the validity of newly proposed species of planktonic foraminifera. Over-splitting of taxa has been a classic problem in foraminiferal research, and studies such as this are necessary to prevent such over-splitting.


From the research team of Dr. Kristian Fauchald and undergraduate student Abigail Knee: 'A cladistic analysis of the polychaete genus Halosydna'. This research helps systematists to clarify the often-problematic evolutionary relationships among a group of marine scale worms. Two new species descriptions contribute to zoologists' understanding of the diversity of animals.


From the research team of Dr. Thomas A. Munroe and undergraduate student participant Trevor J. Krabbenhoft: 'A New Cynoglossid Flatfish (Symphurus: Cynoglossidae: Pleuronectiformes) from Deep-waters of the Philippine Archipelago'. The description of a new species to science contributes to biodiversity studies and conservation efforts by providing information for other scientists to better understand the unique organisms in specific areas and habitats.


From the Research Team of Dr. J. Daniel Rogers and undergraduate student Courtney Porreca: 'Paste Composition and Vessel Function: A Case Study of Caribbean Ceramics'. This research contributes to the knowledge and understanding of Caribbean prehistory by providing preliminary information on technological continuity on an individual island as well as introducing an alternate method for examining ceramic paste composition.


From the research team of Dr. Melinda Zeder and undergraduate student participant Anastasia Poulos: 'The Hunting of Gazella subgutturosa by Early Domesticators in the Fertile Crescent- A Comparison of Wild and Domestic Animal Remains in the Archaeological Record'. This research further reinforces the feasibility of obtaining a pattern for domestication that can be determined empirically. Since domestication has often been interpreted in the archaeological record solely on changes in obvious physical characteristics, such as size change, there has been a long period of time at the beginning of animal domestication, before these characteristic changes took place, which has not been fully recognized. This technique allows for an earlier recognition of domestic remains, while also sheds light on the exploitation of other resources, such as the gazelle.


From the research team of Dr. Carole Baldwin, Dr. Lynne Parenti, Dr. Springer and undergraduate student Julianne Rowehl: 'Identification of Galapagos Entomacrodus (Teleostei: Blennidae) with comments on relationships within the Entomacrodus "nigricans" species group'. This research extended the range of Entomacrodus chiostictus to the Galapagos Archipelago. A preliminary understanding of the biogeography of this fish was established. The evolutions of fish often follow patterns of other similarly distributed fish. An understanding of one fish's evolution could lead to the understanding of many other fish genera.


From the research team of Dr. Paul M. Peterson and undergraduate student Jeffery M. Saarela: 'The taxonomy of Brachyelytrum (Poaceae).' The revisionary study of a group of plants provides a better understanding of the number of distinct species in that group and how to correctly identify them. This information is critical in understanding the biodiversity of our world and to other scientists working with these organisms.


From the Research team of DR. Richard Thorington and undergraduate student Shannon Schwaller: 'A morphometric analysis of hand anatomy in five genera of New World monkeys.' The statistical analysis of hand anatomy provides an important body of knowledge on the development and evolution of the New World primate hand. The quantification of hand anatomy in the sub-family Atelinae contributes important information concerning the underlying genetic processes involved with specific morphological changes in primate hands.


From the research team of Dr. Steven Lingafelter and undergraduate student participant Ainsley Seago: 'A New Species of Crepidodera (Coleoptera: Chrysomelidae)'. This project offers a thorough description of C. praestigiator as well as the means for distinguishing this species from others in the genus Crepidodera. Illustrations produced as part of this project may serve to elucidate the species-specific structure of genitalia among Crepidodera. Description of this new species also contributes to biodiversity studies and knowledge of insects native to the D.C. area.


From the research team of Dr. William Melson and undergraduate student Dawn Stodden: 'The Assembly of North America: Using Garnet Zoning to Decipher the Geologic History of the Potomac Terrane'. This study is a preliminary study of the Sykesville Formation of the Potomac Terrane. Studying terrane in Eastern North America can prove to be a difficult task because of the extensive tectonic history of the area. Researching and reporting the geologic evolution of an area will give future researchers of Eastern North America a good foundation for the evolution of the Potomac Terrane, which could potentially help in their research. The microprobe images and data are a potential aid to a better understanding of zoned minerals.


From the research team of Dr. Vicki Funk and undergraduate student Katarina Topalov: 'Evaluation of Plant Diversity on the Guiana Shield.' The research provides a new approach in overcoming the collecting bias such as time of year or intensity of collecting by using various kinds of statistical analyses. This would provide a more objective application of the already existing data and, therefore, more precise insight into the biodiversity of the explored area.


From the research team of Dr. Ed Vicenzi and undergraduate student Sharon Wilson: 'Microscopy and microchemistry of low temperature aqueous alteration in the Lafayette meteorite.' Lafayette is a Martian meteorite that contains clues about the geologic history of Mars. The meteorite contains rust-like veins that were formed by Martian water. The physical and chemical characteristics of the veins provide crucial information to determine whether the environment beneath the surface of Mars was once capable of sustaining microbial lifeforms.


2. Contributions to other disciplines of science or engineering

From the research team of Dr. Sorena S. Sorensen and undergraduate student Erin DiMaggio: 'Evidence for Feldspar Replacement in Altered Metamorphic Tuffs in the Duck Lake Volcanic Suite'. This research may eventually increase our understanding of ore deposit formation, tectonic history and the geochemical evolution of continental crust during magmatism.


From the research team of Dr. James C. Tyler, Dr. G. David Johnson and undergraduate student Matthew Friedman: 'A putative lampridiform from the Oligocene of Peru'. This research serves as a model to encourage exploration outside of one's established field in order produce more thorough work. Clearly this fossil could have been studied in a purely paleontological sense, but interaction with modern ichthyologists and extensive examination of extant fishes made the research more critical and meaningful.


From the research team of Dr. John Pandolfi and Dr. Ian Macintyre and undergraduate student participant Michael Holcomb: "Use of X-radiographs for coral identification". Many of the characteristics examined are ones that would potentially be available in core samples. Thus this work may lead to better identification of materials within core samples and a consequent improvement in our understanding of past climatic and oceanic conditions. By being able to identify specimens, errors due to vital effects can be reduced, this is of particular importance to geochemists.


From the research team of Dr. Brian Huber and recent graduate Laura Holladay: 'Ontogenetic morphometric comparison of middle-late Eocene pseudohastigerinid planktonic foraminifera using X-radiograph image analysis'. This research provides a model for other disciplines to consider alternative methods of analysis. The use of x-radiography in foraminiferal research is relatively recent, but it has proven to be an ideal method for imaging internal structures. Researchers in other disciplines should also recognize the potential utility of such methods for taxonimic and paleoenvironmental interpretations.


From the research team of Dr. Kristian Fauchald and undergraduate student Abigail Knee: 'A cladistic analysis of the polychaete genus Halosydna'. This research can be used as a part of a study on cladistic methodology. It also provides clues to evolutionary biologists with the example of 18 species in question.


From the Research Team of Dr. J. Daniel Rogers and undergraduate student Courtney Porreca: 'Paste Composition and Vessel Function: A Case Study of Caribbean Ceramics'. The information regarding technological, and possibly cultural continuity is important in the study of cultural traditions, culture contact, and cultural change.


From the research team of Dr. Melinda Zeder and undergraduate student participant Anastasia Poulos: 'The Hunting of Gazella subgutturosa by Early Domesticators in the Fertile Crescent- A Comparison of Wild and Domestic Animal Remains in the Archaeological Record'. This project has not had a solely archaeological impact, since it also focused on a collection of modern gazelle. A number of biological implications came forward under this study, such as strong regional variation in the gazelle of the area today.


From the research team of Dr. Carole Baldwin, Dr. Lynne Parenti, Dr. Springer and undergraduate student Julianne Rowehl: 'Identification of Galapagos Entomacrodus (Teleostei: Blennidae) with comments on relationships within the Entomacrodus "nigricans" species group'. This research may be used to support theories of continental drift in relation to biogeography by giving examples of evolution based on plate movement.


From the Research team of Dr. Richard Thorington and undergraduate student Shannon Schwaller: 'A morphometric analysis of hand anatomy in five genera of New World monkeys'. This research builds upon a body of knowledge concerning primate growth and development that could be useful to medical and molecular. The results provided in this study could contribute valuable information to genetic researchers that are interested in effects of genetic change and development of the hand.


From the research team of Dr. Steven Lingafelter and undergraduate student participant Ainsley Seago: 'A New Species of Crepidodera (Coleoptera: Chrysomelidae)'. As the family Chrysomelidae includes many agricultural pests, the broader knowledge of chrysomelid diversity created by this research applies to the economics and knowledge-based counteraction of insect damage to crops and ornamental plants.


From the research team of Dr. William Melson and undergraduate student Dawn Stodden: 'The Assembly of North America: Using Garnet Zoning to Decipher the Geologic History of the Potomac Terrane'. Geophysicists can use the research we have done to assist them with similar geophysical perplexities.


From the research team of Dr. Ed Vicenzi and undergraduate student Sharon Wilson: 'Microscopy and microchemistry of low temperature aqueous alteration in the Lafayette meteorite.' The search for life on Mars extends into the field of biology to find a terrestrial analogue for low-temperature aqueous environments that are capable of sustaining microbial lifeforms.


3. Contributions to the development of human resources

From the research team of Dr. Sorena S. Sorensen and undergraduate student Erin DiMaggio: 'Evidence for Feldspar Replacement in Altered Metamorphic Tuffs in the Duck Lake Volcanic Suite'. The results of this research produced an abstract that was submitted to the Geological Society of America for the annual meeting in November of 2001 and will be presented as a talk. The abstract and the talk are available to the public, a sharing of ideas that will serve as a stimulant for further support of research.


From the research team of Dr. James C. Tyler, Dr. G. David Johnson and undergraduate student Matthew Friedman: 'A putative lampridiform from the Oligocene of Peru'. The findings of this study will be presented to high school students and teachers, as well as non-science oriented college students, and will hopefully work to engender science and scientific research to those audiences.


From the research team of Dr. John Pandolfi and Dr. Ian Macintyre and undergraduate student participant Michael Holcomb: "Use of X-radiographs for coral identification". Through presenting this research to undergraduate students, non-scientists and future pre-college teachers may be exposed to additional fields to which they might otherwise not have heard about.


From the research team of Dr. Kristian Fauchald and undergraduate student Abigail Knee: 'A cladistic analysis of the polychaete genus Halosydna'. This research provided a female undergraduate with the opportunity to learn the value and techniques of collection-based research in systematics, which will be an important experience as she goes on to graduate studies and a career in research.


From the research team of Dr. Carole Baldwin, Dr. Lynne Parenti, Dr. Springer and undergraduate student Julianne Rowehl: 'Identification of Galapagos Entomacrodus (Teleostei: Blennidae) with comments on relationships within the Entomacrodus "nigricans" species group'. Support of maintained collections, such as the fish collection at the NMNH, is invaluable when trying to understand relationships and biodiversity.


From the research team of Dr. Steven Lingafelter and undergraduate student participant Ainsley Seago: 'A New Species of Crepidodera (Coleoptera: Chrysomelidae)'. The collections and conclusions resulting from this research will be useful in any future investigations (through research, as part of an educational process, etc) of chrysomelid and/or alticine phylogeny and morphology.


4. Contributions to the physical, institutional, or information resources that form the infrastructure for research and education

From the research team of Dr. Sorena S. Sorensen and undergraduate student Erin DiMaggio: 'Evidence for Feldspar Replacement in Altered Metamorphic Tuffs in the Duck Lake Volcanic Suite'. This project, in addition to the databases created, can serve as a systematic way of looking at rocks from this area and can be followed by future research scientists who are studying alteration patterns and effects of hydrothermal systems.


From the research team of Dr. James C. Tyler, Dr. G. David Johnson and undergraduate student Matthew Friedman: 'A putative lampridiform from the Oligocene of Peru'. This project serves as a model for future researchers and encourages them to examine fossils as biota and in the context of related extant taxa. Such an approach has potential for more thorough research in the life sciences, deconstructing the artifactual division between paleontology and modern organismal biology.


From the research team of Dr. Kristian Fauchald and undergraduate student Abigail Knee: 'A cladistic analysis of the polychaete genus Halosydna'. The data collected from this research will hopefully contribute to a future review of the polychaete family Polynoidae, giving clues to evolutionary significant characters.


From the Research Team of Dr. J. Daniel Rogers and undergraduate student Courtney Porreca: 'Paste Composition and Vessel Function: A Case Study of Caribbean Ceramics'. This research utilizes analytical methods that do not require the use of highly technical equipment, providing an alternate to more traditional methods and allowing for easier access to this type of research.


From the research team of Dr. Melinda Zeder and undergraduate student participant Anastasia Poulos: 'The Hunting of Gazella subgutturosa by Early Domesticators in the Fertile Crescent- A Comparison of Wild and Domestic Animal Remains in the Archaeological Record'. This project has made people more aware of the potential importance of zooarchaeological remains, as well as extensive modern collections in being highly educational in regards to human exploitation of their natural resources.


From the research team of Dr. Carole Baldwin, Dr. Lynne Parenti, Dr. Springer and undergraduate student Julianne Rowehl: 'Identification of Galapagos Entomacrodus (Teleostei: Blennidae) with comments on relationships within the Entomacrodus "nigricans" species group'. This research demonstrates the need for collections in tropical areas. Much of the work performed would not have been possible without travel to or research stations developed in these areas. This project can serve as a preliminary work for future research teams.


From the research team of Dr. William Melson and undergraduate student Dawn Stodden: 'The Assembly of North America: Using Garnet Zoning to Decipher the Geologic History of the Potomac Terrane'. Geophysicists can use the research we have done to assist them with similar geophysical perplexities.


From the research team of Dr. Ed Vicenzi and undergraduate student Sharon Wilson: 'Microscopy and microchemistry of low temperature aqueous alteration in the Lafayette meteorite.' The results from this study are reproducible, a valuable and desired aspect in scientific research. The method of data collection serves as an example for future technological studies on Lafayette and similar Martian meteorites.


5. Contributions to other aspects of public welfare beyond science and engineering, such as commercial technology, the economy, cost-efficient environmental protection, or solutions to social problems.

From the research team of Dr. Sorena S. Sorensen and undergraduate student Erin DiMaggio: 'Evidence for Feldspar Replacement in Altered Metamorphic Tuffs in the Duck Lake Volcanic Suite'. Studying fluid flow through rocks has important practical economic value. This research may help in understanding ore deposit formation which is an important process to interpret due to its economic significance.


From the research team of Dr. James C. Tyler, Dr. G. David Johnson and undergraduate student Matthew Friedman: 'A putative lampridiform from the Oligocene of Peru'. Though much of the research performed was paleontological in nature, the project still had strong groundings in biology. Like other biological research, this project highlights the diversity of the biotic world, both Recent and extinct, and stresses the need to preserve current biodiversity.


From the Research Team of Dr. J. Daniel Rogers and undergraduate student Courtney Porreca: 'Paste Composition and Vessel Function: A Case Study of Caribbean Ceramics'. The research resulted in information about the prehistory of a Caribbean island, which could be useful in promoting tourism.




PERSONAL SUMMARIES FROM OUR SUMMER



Yolanda Chacon

My goals for the RTP included learning the methods for conducting museum-based research on dry specimens of birds and becoming aware of the diversity of research that goes on behind the scenes at the Smithsonian Institution, National Museum of Natural History. Although I am not very familiar with the many aspects of natural history, I applied to the program knowing that by becoming accepted, I would broaden my knowledge, especially in Ornithology. My advisor, Dr. Carla J. Dove proposed the topic of our RTP project, Analysis of feather characters of the Cranes, Rails and Allies (Avian Order:Gruiformes). From the first day of the RTP program, I was very eager to get my project underway and learn the techniques that will help me continue this study at my home institution (New Mexico State University) under the direction of Dr. Peter Houde. My advisor, along with Dr. Peter Houde, suggested the species for use in our study and I investigated the published accounts of this particular order of birds, looking for traditional phylogenic analysis and those obtained from DNA characters. With the assistance of my advisor, and Marcy Heacker-Skeans (a master's student at George Mason University) we described, photographed and investigated (using parsimony analysis) 12 species within the order Gruiformes and one outgroup (Galliformes) to determine if feather characters can provide phylogenetic signal. Suzanna Braden of the NMNH Scanning Electron Microscope (SEM) laboratory assisted the SEM work. Microscopic feather character coding was done at NMNH Division of Birds under the assistance of my project advisor. From this study, we hope to publish in summer 2002, and as stated previously, I will continue this study at New Mexico State University.


Other things I did during the summer

1. Behind-the-scenes tour of the NMNH for SERC, CRC and ZOO interns. Hosted the ornithology section of the tour and presented a 10 minute demonstration on the Feather Identification Lab and my RTP research topic, Analysis of feather characters of the Cranes Rails and Allies from (Order: Gruiformes)

2. Poster stuffing pizza party. Participant in the RTP poster stuffing pizza party, prepared 7,000 2002 RTP advertisement posters.

3. Behind-the-scenes tour of the NMNH for fellow RTP interns. Served as an organizer and host The Bird Collection, Museum Support Center- Dry Mammals Storage, Osteo Prep and The New Whale Storage Facility. Recruited 18 student participants and confirmed tour presenters.

4. The Brain. (13 July 2001). Attended a Symposium held at the Smithsonian Institution NMNH Baird Auditorium.

5. Bird Watching. Participated in bird watching with Marcy Heaker-Skeans to better familiarize myself with the local bird diversity and learn field identification skills.


Erin DiMaggio

Extensive studies on the alteration histories of volcanic rocks in eastern Sierra Nevada have been done by SI researchers. Although scientists know what rocks surround Duck Lake, until now no one had looked at the petrography of this group of metamorphic tuffs. The purpose of my project was to examine and analyze feldspar textures and mineral chemistry in order to determine the processes responsible for the alteration of these rocks. In addition, my results were compared with the chemical findings of the surrounding areas in order to determine how the geologic history of rocks in this area differed. In order to accomplish this three main instruments were used. First, and used most frequently, was cathodoluminescence (CL). I used CL to determine the growth and replacement history of feldspars in these rocks. With instrument assistance from my research advisor, Sorena Sorensen and Ed Vicenzi the CL proved to be a useful tool in identifying alteration patterns. SEM and Microprobe work was aided by Amelia Logan and James Rougvie. Using backscattered electron imaging and EDS analyses on the SEM I was able to qualitatively determine mineral composition in order to validate my CL observations and obtain images that would aid in navigating the microprobe. An electron microprobe was used to quantitatively determine mineral compositions and produce element X-ray composition maps. The X-ray maps were then used to analyze the location and distribution of major elements and were compared with CL and backscattered images. A scientific abstract was produced in collaboration with James Rougvie and Sorena Sorensen and submitted to the Geological Society of America for the 2001 annual meeting. Sorensen and Rougvie will continue work on the tuffs after the conclusion of this internship. In addition, my research will be incorporated in a larger scale study of the area and later published.

Other things I did during the summer

1. Great North ( 22 June 2001 ). Viewed and critiqued a new IMAX film at the NMNH.

2. Behind-the-scenes tour of the NMNH for CRC and SERC interns. (6th July 2001) Hosted a tour of the Meteorites given by Tim McCoy, the curator of the Meteorites.

3. Tour of the Capital led by an intern working for Senator Susan Collins
of Maine. (29th June 2001) I was told the history and little secrets of the Capital building and I attended the Senate in session.

4. Tour of the Gems and Minerals Collection given by Jeff Post (26th June 2001) Viewed new additions and old favorites in the Gems and Minerals Collection.

5. Department of Mineral Sciences Lecture Series: Dick Fiske and Tim Rose "Kulanaokuaiki 3 Tephra: Kilauea Volcano's Nasty Little Secret" (21st June 2001) Highlighting one of Hawaii's gentle volcanoes, ongoing research is pointing towards a new and rather unusual direction. Could this quiet volcano actually be the culprit of launching large clasts more than 6 km from the summit?

6. Department of Mineral Sciences Lecture Series: Ed Vicenzi "On the Origin of Precambrian Black Diamond (Carbonado) from Central Africa and Brazil: Constraints from Microanalytical Studies" (20th July 2001) Several hypothesis have been proposed as to the origin of such fused diamond crystallites. Each proposal could play a crucial role in our understanding of early Earth.

7. Smithsonian Lecture Series: Dr. Robert T. Watson "The Perils of Ignoring Global Warming: State of Science and Politics" (15th June 2001) Dr. Watson presented information about the current political situation of the Kyoto Protocol in addition to a brief over-view and summary of the recent Third Assessment Report.

8. Attended the viewing of Cirque du Soleil an IMAX film presented by the NMNH followed by the Friday night Jazz Café (15th & 29th June 2001).

9. Smithsonian Folklife Festival (29th & 30th June, 6th July 2001) Attended this spectacular display of the components that make each cultural so truly unique including food, dance and music.


Matthew Scott Friedman

My childhood exposure to some of the finest Devonian fish fossils ever found, collected from outcrops fifteen minutes away from my home, led to an early interest in paleoichthyology. I had hoped that my experience in the RTP would give me valuable experience in the basics of examining and describing fossil fish, but my work at the NMNH has exceeded those initial expectations. With most of my background in Paleozoic fishes, I found being confronted with an Oligocene teleost by my advisors (Dr. JAMES C. TYLER and Dr. G. DAVID JOHNSON; the fossil was brought to Dr. Tyler's attention by key contact BOB PURDY) particularly interesting and challenging. The relatively young fossil allowed for comparison with specimens of Recent taxa, giving me my fist real opportunity to examine extant bony fishes in detail. When I compared the fossil to the living members of the family Luvaroidea, the fishes to which Dr. Tyler thought the specimen was most closely allied, I found numerous inconsistencies and disparities in osteology. Working together with my two advisors, we concluded that the fish was more likely a member of the order Lampridiformes, represented in the Recent fauna by such unusual fishes as the opah and the oarfish. In order to examine obscured osteological characters for comparative work, CT scan data were taken with the assistance of EVAN GAROFALO of the Anthropology Department using Dr. BRUNO FROHLICH's machine. More detailed examinations were made possible through the use of more conventional preparation techniques, such as acid treatment and mechanical abrasion. I was instructed in these techniques by FRED GRADY, a preparator in the Vertebrate Paleontology Preparation Lab. I spent several weeks slowly grinding away the siliceous matrix that obscured distinctive or potentially diagnostic characters with a diamond-tipped burr, removing larger pieces of matrix with a diamond saw. Following preparation, Dr. Tyler and I described the osteology of the specimen. Afterwards, I compared the anatomy of the fossil skull to that observed in dried, alcohol-preserved, and cleared and stained specimens of lampridiform fish. Dr. JOHN G. MAISEY of the American Museum of Natural History and Dr. GLORIA ARATTIA of Humbolt University offered advice and perspectives on examining the fossil, while Dr. JOHN OLNEY of the Virginia Institute of Marine Science provided helpful images. Cleared and stained specimens of higher lampridiform taxa were generously loaned by the Scripps Institute of Oceanography, and were delivered to us at the ASIH meetings by Dr. H. J. WALKER and Dr. C. KLEPADLO. BARBARA BROWN, MELANIE STIASSNY, RADFORD ARRINDELL and IVY RUTSKY provided invaluable assistance in procuring specimens from the American Museum of Natural History. Possibilities of more detailed CT scan data were discussed with TIM ROWE of the University of Texas. Though the project will not be completed by the end of the RTP session, we intend to continue researching this interesting specimen with the intention of publishing a paper detailing our results. Ideally, I would like to work on this project extensively at school, and allow it to develop into a senior thesis. My research on this fossil in concert with specimens of extant fishes has kindled an interest in me to study extinct and living taxa in unison to more completely understand the evolutionary history of clades. I may pursue research on living and fossil representatives of the order Lampridiformes in graduate school.

Other things I did during the summer

1. American Museum of Natural History (8 June, 23 June). Attended AMNH on two separate occasions during my stay in Washington, D.C. On the first trip, my advisor and I went to the museum to pick up specimens for comparative work, and he took the opportunity to introduce me to several of the scientists working there. On the second trip, four other RTP interns and myself visited AMNH in order to tour the exhibits more extensively.

2. Invertebrate Zoology Seminar (28 June). Attended a lecture regarding weevil diversity and classification given by a scientist visiting from the Natural History Museum (London).

3. National Aquarium in Baltimore (1 July). Four other RTP interns and I attended the aquarium in order to view the animals on display.

4. ASIH/AES Joint Meetings (6-10 July). Attended several lectures given by both societies as well as ASIH social events. Spoke to several scientists about graduate school possibilities.

5. Paleobiology Seminar (19 July). Attended a lecture with other RTP interns regarding the evolution of whales based on fossils.

6. National Zoo (22 July). Attended the National Zoological Park to observe the animals on display.

7. Vertebrate Zoology Seminar (24 July). Attended a lecture with other RTP interns about fossil flatfishes.

8. Independent Biology Research. Spoke with several Smithsonian scientists about potential groups to use for a paper a professor at the University of Rochester and I are writing about the reconstruction of phylogenies using morphological, stratigraphic, and molecular data for publication in Paleobiology.

9. Independent Paleontology Research. Spoke with paleobotanist Amanda Ash regarding the possibility of identification of fossils collected on a research expedition to the Canadian arctic that I took part in. Examined the vertebrate paleontology collections in an attempt to identify teeth and other elements collected in the arctic to the family or genus level.


Michael Holcomb

During the course of this project I digitized approx. 100 X-ray images of coral slabs. I took measurements on these images and on images that had been previously digitized. I placed my measurement values into a database and then conducted statistical analysis on the data sets to determine their potential for use in distinguishing members of the Montastraea annularis species complex. This project was an extension of work being conducted by my advisor, Dr. John Pandolfi in collaboration with Dr. Ann F. Budd. This work also marked further developments and applications of methods developed some two decades ago by my advisor Dr. Ian Macintyre. In the course of my project, I was assisted by Bill Boykins with X-rays, and by Dr. Lee-Ann Hayek with statistics. Though this particular aspect of the work is not something I will be continuing, I will follow up this work by doing some more in depth elemental analysis, looking for possible vital effects upon composition and localization of skeletal elements.

Other things I did during the summer

1. North American Paleontological Conference (June 26 - June 30, 2001). Attended various lectures and other presentations.

2. Attended various seminars on topics ranging from the evolution of whales, to Phragmites australis and its invasion of the North East, to coral paleoecology (May 28 - July 26, 2001).


Laura Holladay

My undergraduate degree is in geological sciences, and my previous research experience fell within the field of geomagnetism. In graduate school and beyond, however, I plan to focus on paleoclimate research. I saw the RTP program as an opportunity to learn more about paleobiology, particularly of the microscopic foraminifera that are so often utilized in paleoclimatic studies. Although the majority of former RTP students who have worked under my advisor (Dr. BRIAN HUBER) have conducted projects focusing on isotopic studies of foraminifera, I wanted to take advantage of the opportunity to study foraminifera as biological entities rather than merely as geochemical tools. I expressed the desire to pursue a project focusing on the morphology and paleobiology of foraminifera, and Dr. Huber suggested a project focusing on the ontogenetic morphometrics of four similar planispiral morphogroups of planktonic foraminifera from the Eocene. Detailed morphometric analyses of these four groups would reveal whether they should be classified as distinct species, and a follow-up study of their isotopic compositions would reveal information about their preferred depth habitats within the water column when they were alive. The specimens chosen for measurement had been prepared and named by Dr. RICHARD OLSSON of Rutgers University, who also provided SEM images to supplement the study. I mounted these specimens onto slides and then produced and developed X-radiograph images, which revealed the microfossils' inner structures. The X-ray films were then mounted on separate slides so that they could be examined with a transmitted-light microscope. I obtained digital images of each specimen X-ray by using a video camera mounted onto the microscope. Measurements were taken from these digital images using ImagePro biometric software. In addition to discrete length and area measurements, RALPH CHAPMAN assisted the morphometric data gathering by providing a computer program that calculates the logarithmic spiral traced out by the ontogenetic addition of chambers. Additional samples were then selected for isotopic study. I prepared the samples with the assistance of MY LE DUCHARME, and Dr. Huber assisted the picking of individual specimens. The specimens were then sent to Dr. KYGER C. LOHMANN of the University of Michigan for carbon and oxygen isotopic analysis. With my project advisor, I analyzed the morphometric data collected. The results were prepared into lecture and poster formats for presentation to the Smithsonian Institution community. My advisor and I are both interested in continuing this and other similar research projects in the future, and the final report for this program will be submitted for publication in The Journal of Foraminiferal Research.

Other things I did during the summer

1. Behind-the-scenes tour of the NMNH for SERC and ZOO interns. Hosted the paleontology tour and presented a 10 minute demonstration on the use of X-ray imaging for foraminiferal research.

2. Natural History Film and Lecture Series. Attended Friday noon lectures and films on topics ranging from archeology to global climate change.

3. The Great North. Attended a special staff preview of a new IMAX film.

4. IMAX Jazz Café. Attended a film and music series that took place at the NMNH after museum hours on Friday nights.

5. Smithsonian Institution Folklife Festival. Visited the concerts, educational demonstrations, and exhibits of this annual multicultural celebration.


Abigail Knee

I have loved the ocean since I was a little girl growing up near the New Hampshire coast. The diverse forms and interactions of marine invertebrates especially captivate me. To complement my undergraduate education in marine biology, special interests in ecology and systematics, and experience in field research, my goals for the RTP program were to learn how collection-based systematists classify species and construct phylogenies. These are skills I would like to apply to my career plans for exploratory research of deep-sea invertebrate communities. In the NMNH Division of Worms, my vehicle for such systematic research was a genus of marine scale worms, under the direction of my project advisor, Dr. KRISTIAN FAUCHALD. The entire class of Polychaeta is full of problematic classification, therefore requiring constant revision as new data becomes available. My advisor is undertaking a long-term project of reconstructing the phylogeny of the polychaete family Polynoidae, among others, and the genus I proposed to work on fits into his research. In my project, I have examined mainly external morphological characters of ten species of Halosydna in the NMNH collection for a cladistic analysis. Using my data on Halosydna spp. and my project advisor's data on eight sister taxa he scored previously, I have reconstructed a series of cladograms using the software program PAUP. Kristian pointed me towards interpretation of the scale worm phylogeny in question. MICHAEL GUTKNECHT taught me how to search the literature for additional background information. LINDA WARD was also available for suggestions on how to report my research. Both Michael and Linda provided technical assistance with equipment. JOANA SILVA, a participant in the RTP '99 under my advisor, was instrumental in instructing me about systematic methods she has employed fo